Institutional 13F holdings sum to 100.4% of shares outstanding. Readings above 100% reflect securities-lending double counting: shares on loan to short sellers are reported long by both the lending institution and the one that bought them. FINRA short interest of 16,117,209 shares (settlement 2026-04-30) accounts for the excess.
| # | Holder | Shares | Value | % of shares out | Δ QoQ |
|---|---|---|---|---|---|
| 1 | BlackRock, Inc. | 17,998,32918.0M | $161M | 12.90% | ADD 10% |
| 2 | ARCH Venture Management, LLC | 12,916,66312.9M | $116M | 9.26% | — |
| 3 | SB INVESTMENT ADVISERS (UK) LTD | 10,948,09310.9M | $98M | 7.85% | TRIM -17% |
| 4 | VANGUARD PORTFOLIO MANAGEMENT LLC | 9,215,5779.2M | $83M | 6.61% | NEW† |
| 5 | GSK plc | 8,550,9548.6M | $77M | 6.13% | — |
| 6 | STATE STREET CORP | 7,141,0947.1M | $64M | 5.12% | ADD 15% |
| 7 | VANGUARD CAPITAL MANAGEMENT LLC | 5,382,1035.4M | $48M | 3.86% | NEW† |
| 8 | GOLDMAN SACHS GROUP INC | 3,250,2273.3M | $29M | 2.33% | ADD 107% |
| 9 | MORGAN STANLEY | 2,854,2052.9M | $26M | 2.05% | ADD 7% |
| 10 | ADAR1 Capital Management, LLC | 2,744,7142.7M | $25M | 1.97% | NEW |
| 11 | GEODE CAPITAL MANAGEMENT, LLC | 2,657,8432.7M | $24M | 1.91% | ADD 14% |
| 12 | DIMENSIONAL FUND ADVISORS LP | 2,579,6782.6M | $23M | 1.85% | — |
| 13 | CITADEL ADVISORS LLC | 2,098,4762.1M | $19M | 1.50% | ADD 329% |
| 14 | FMR LLC | 2,071,8482.1M | $19M | 1.49% | TRIM -34% |
| 15 | Soleus Capital Management, L.P. | 2,002,0002.0M | $18M | 1.43% | NEW |
| 16 | Point72 Asset Management, L.P. | 1,952,2342.0M | $17M | 1.40% | ADD 897% |
| 17 | Boxer Capital Management, LLC | 1,866,0001.9M | $17M | 1.34% | ADD 199% |
| 18 | Avidity Partners Management LP | 1,569,3001.6M | $15M | 1.12% | NEW |
| 19 | GordonMD Global Investments LP | 1,518,5521.5M | $14M | 1.09% | ADD 63% |
| 20 | Sio Capital Management, LLC | 1,498,6201.5M | $13M | 1.07% | ADD 213% |
| 21 | Pictet Asset Management Holding SA | 1,480,8281.5M | $13M | 1.06% | ADD 10879% |
| 22 | Hudson Bay Capital Management LP | 1,469,5101.5M | $13M | 1.05% | ADD 11% |
| 23 | JACOBS LEVY EQUITY MANAGEMENT, INC | 1,442,3361.4M | $13M | 1.03% | NEW |
| 24 | TWO SIGMA INVESTMENTS, LP | 1,439,4941.4M | $13M | 1.03% | ADD 144% |
| 25 | Qube Research & Technologies Ltd | 1,248,6691.2M | $11M | 0.89% | ADD 1403% |
| 26 | Clearstead Advisors, LLC | 1,216,4001.2M | $11M | 0.87% | NEW |
| 27 | Jones Hill Capital LP | 1,198,2131.2M | $11M | 0.86% | NEW |
| 28 | CHARLES SCHWAB INVESTMENT MANAGEMENT INC | 1,171,6841.2M | $10M | 0.84% | ADD 30% |
| 29 | Candriam S.C.A. | 1,135,7631.1M | $10M | 0.81% | ADD 10% |
| 30 | BNP PARIBAS FINANCIAL MARKETS | 1,078,0201.1M | $10M | 0.77% | ADD 285% |
| 31 | Ensign Peak Advisors, Inc | 1,070,8121.1M | $10M | 0.77% | ADD 101% |
| 32 | JPMORGAN CHASE & CO | 1,117,5161.1M | $9M | 0.80% | ADD 167% |
| 33 | Nuveen, LLC | 1,018,9641.0M | $9M | 0.73% | ADD 80% |
| 34 | NORTHERN TRUST CORP | 985,435985,435 | $9M | 0.71% | ADD 16% |
| 35 | BANK OF AMERICA CORP /DE/ | 965,814965,814 | $9M | 0.69% | ADD 4% |
| 36 | D. E. Shaw & Co., Inc. | 817,685817,685 | $7M | 0.59% | ADD 1542% |
| 37 | RENAISSANCE TECHNOLOGIES LLC | 761,364761,364 | $7M | 0.55% | ADD 24% |
| 38 | UBS Group AG | 744,142744,142 | $7M | 0.53% | ADD 78% |
| 39 | VANGUARD FIDUCIARY TRUST CO | 731,721731,721 | $7M | 0.52% | NEW |
| 40 | SUPERSTRING CAPITAL MANAGEMENT LP | 730,548730,548 | $7M | 0.52% | NEW |
| 41 | Walleye Capital LLC | 665,584665,584 | $6M | 0.48% | ADD 160% |
| 42 | Bank of New York Mellon Corp | 584,047584,047 | $5M | 0.42% | ADD 7% |
| 43 | NORDEA INVESTMENT MANAGEMENT AB | 580,755580,755 | $5M | 0.42% | TRIM -5% |
| 44 | AQR CAPITAL MANAGEMENT LLC | 566,009566,009 | $5M | 0.41% | ADD 204% |
| 45 | PRINCIPAL FINANCIAL GROUP INC | 547,753547,753 | $5M | 0.39% | ADD 22% |
| 46 | Empyrean Capital Partners, LP | 527,800527,800 | $5M | 0.38% | NEW |
| 47 | Alta Partners Management Company, L.P. | 509,917509,917 | $5M | 0.37% | — |
| 48 | Parkman Healthcare Partners LLC | 500,000500,000 | $4M | 0.36% | NEW |
| 49 | MILLENNIUM MANAGEMENT LLC | 489,360489,360 | $4M | 0.35% | TRIM -51% |
| 50 | ALGERT GLOBAL LLC | 489,008489,008 | $4M | 0.35% | NEW |
| 51 | Invesco Ltd. | 421,332421,332 | $4M | 0.30% | ADD 4% |
| 52 | Public Sector Pension Investment Board | 403,795403,795 | $4M | 0.29% | ADD 24% |
| 53 | AMERIPRISE FINANCIAL INC | 395,038395,038 | $4M | 0.28% | TRIM -46% |
| 54 | PANAGORA ASSET MANAGEMENT INC | 375,354375,354 | $3M | 0.27% | — |
| 55 | Schonfeld Strategic Advisors LLC | 370,128370,128 | $3M | 0.27% | NEW |
| 56 | WINTON GROUP Ltd | 367,883367,883 | $3M | 0.26% | ADD 94% |
| 57 | Susquehanna Portfolio Strategies, LLC | 353,663353,663 | $3M | 0.25% | ADD 646% |
| 58 | AMERICAN CENTURY COMPANIES INC | 296,824296,824 | $3M | 0.21% | — |
| 59 | Corient Private Wealth LLC | 293,688293,688 | $3M | 0.21% | TRIM -44% |
| 60 | Ikarian Capital, LLC | 293,258293,258 | $3M | 0.21% | TRIM -49% |
| 61 | Legal & General Group Plc | 256,749256,749 | $2M | 0.18% | ADD 7% |
| 62 | CITIGROUP INC | 256,140256,140 | $2M | 0.18% | TRIM -18% |
| 63 | Engineers Gate Manager LP | 250,391250,391 | $2M | 0.18% | NEW |
| 64 | Eversept Partners, LP | 250,000250,000 | $2M | 0.18% | NEW |
| 65 | RHUMBLINE ADVISERS | 232,550232,550 | $2M | 0.17% | ADD 12% |
| 66 | Swiss National Bank | 231,500231,500 | $2M | 0.17% | ADD 30% |
| 67 | SUSQUEHANNA INTERNATIONAL GROUP, LLP | 195,561195,561 | $2M | 0.14% | TRIM -50% |
| 68 | Man Group plc | 177,924177,924 | $2M | 0.13% | NEW |
| 69 | Erste Asset Management GmbH | 180,800180,800 | $2M | 0.13% | ADD 528% |
| 70 | Tejara Capital Ltd | 161,789161,789 | $1M | 0.12% | TRIM -30% |
| 71 | Hillsdale Investment Management Inc. | 161,096161,096 | $1M | 0.12% | NEW |
| 72 | ExodusPoint Capital Management, LP | 137,059137,059 | $1M | 0.10% | TRIM -23% |
| 73 | BARCLAYS PLC | 134,796134,796 | $1M | 0.10% | TRIM -63% |
| 74 | Y-Intercept (Hong Kong) Ltd | 134,547134,547 | $1M | 0.10% | NEW |
| 75 | Rafferty Asset Management, LLC | 134,413134,413 | $1M | 0.10% | — |
| 76 | UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC | 130,503130,503 | $1M | 0.09% | ADD 8% |
| 77 | NEW YORK STATE TEACHERS RETIREMENT SYSTEM | 122,900122,900 | $1M | 0.09% | TRIM -3% |
| 78 | Jump Financial, LLC | 122,066122,066 | $1M | 0.09% | NEW |
| 79 | Raiffeisen Bank International AG | 125,000125,000 | $1M | 0.09% | NEW |
| 80 | PRUDENTIAL FINANCIAL INC | 115,402115,402 | $1M | 0.08% | TRIM -59% |
| 81 | Thrivent Financial for Lutherans | 113,593113,593 | $1M | 0.08% | ADD 75% |
| 82 | OP Asset Management Ltd | 113,028113,028 | $1M | 0.08% | NEW |
| 83 | MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd. | 113,338113,338 | $1M | 0.08% | ADD 6% |
| 84 | PUBLIC EMPLOYEES RETIREMENT SYSTEM OF OHIO | 112,700112,700 | $1M | 0.08% | NEW |
| 85 | Headlands Technologies LLC | 103,247103,247 | $925,093 | 0.07% | ADD 358% |
| 86 | Creative Planning | 100,511100,511 | $900,579 | 0.07% | ADD 26% |
| 87 | Monashee Investment Management LLC | 100,000100,000 | $896,000 | 0.07% | NEW |
| 88 | PRICE T ROWE ASSOCIATES INC /MD/ | 96,14596,145 | $862,000 | 0.07% | ADD 18% |
| 89 | IEQ CAPITAL, LLC | 96,01096,010 | $860,249 | 0.07% | NEW |
| 90 | Quantinno Capital Management LP | 95,92795,927 | $859,506 | 0.07% | ADD 144% |
| 91 | Trexquant Investment LP | 94,76794,767 | $849,112 | 0.07% | NEW |
| 92 | THOMPSON SIEGEL & WALMSLEY LLC | 92,00092,000 | $824,000 | 0.07% | NEW |
| 93 | DEUTSCHE BANK AG\ | 90,73390,733 | $812,968 | 0.07% | TRIM -56% |
| 94 | CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM | 87,37587,375 | $782,880 | 0.06% | — |
| 95 | Russell Investments Group, Ltd. | 82,21382,213 | $736,626 | 0.06% | ADD 996% |
| 96 | VANGUARD ASSET MANAGEMENT, Ltd | 76,92876,928 | $689,275 | 0.06% | NEW |
| 97 | GSA CAPITAL PARTNERS LLP | 76,41676,416 | $685,000 | 0.05% | ADD 206% |
| 98 | Quadrature Capital Ltd | 75,65875,658 | $677,896 | 0.05% | ADD 52% |
| 99 | Aquatic Capital Management LLC | 75,30175,301 | $674,697 | 0.05% | NEW |
| 100 | TUDOR INVESTMENT CORP ET AL | 73,50473,504 | $658,596 | 0.05% | ADD 7% |
† a filer this large appearing as NEW usually reflects a renamed or succeeded filing entity (the old name stops filing the same quarter) — not a brand-new position.
Rows rank by the value each filer reported — 13F filers price their own positions, so a holder with fewer shares can rank just above one with more.
| Holder | 2024-Q4 | 2025-Q1 | 2025-Q2 | 2025-Q3 | 2025-Q4 | 2026-Q1 | Latest value |
|---|---|---|---|---|---|---|---|
| BlackRock, Inc. | 16.5M | 16.5M | 16.2M | 16.0M | 16.3M | 18.0M | $161M |
| ARCH Venture Management, LLC | — | — | 12.9M | 12.9M | 12.9M | 12.9M | $116M |
| SB INVESTMENT ADVISERS (UK) LTD | 16.7M | 16.7M | 16.7M | 16.7M | 13.1M | 10.9M | $98M |
| VANGUARD PORTFOLIO MANAGEMENT LLC | — | — | — | — | — | 9.2M | $83M |
| GSK plc | — | — | — | — | 8.6M | 8.6M | $77M |
| STATE STREET CORP | 5.1M | 5.7M | 5.3M | 5.2M | 6.2M | 7.1M | $64M |
| VANGUARD CAPITAL MANAGEMENT LLC | — | — | — | — | — | 5.4M | $48M |
| GOLDMAN SACHS GROUP INC | 517K | 743K | 1.3M | 1.5M | 1.6M | 3.3M | $29M |
| MORGAN STANLEY | 1.1M | 1.4M | 1.6M | 2.7M | 2.7M | 2.9M | $26M |
| ADAR1 Capital Management, LLC | 64K | 250K | 250K | — | — | 2.7M | $25M |
| GEODE CAPITAL MANAGEMENT, LLC | 2.2M | 2.3M | 2.3M | 2.2M | 2.3M | 2.7M | $24M |
| DIMENSIONAL FUND ADVISORS LP | 1.9M | 2.7M | 3.0M | 3.0M | 2.5M | 2.6M | $23M |
| CITADEL ADVISORS LLC | 271K | 1.6M | 3.5M | 799K | 489K | 2.1M | $19M |
| FMR LLC | 7K | 2.2M | 3.6M | 3.6M | 3.2M | 2.1M | $19M |
| Soleus Capital Management, L.P. | — | — | — | — | — | 2.0M | $18M |
Shares reported each quarter by the largest current holders. Green = added vs the prior filing, red = trimmed, grey = roughly held; a dash is a quarter with no 13F on file (a gap is skipped, not read as an exit). 13F is filed up to 45 days after quarter-end. Up to 40 quarters (10 years) available — use the selector above.
Line breaks are honest gaps — quarters with no 13F record, or a suppressed % — never interpolated. % of shares outstanding divides each quarter's as-filed 13F shares by the SEC cover-page share count dated to that same quarter (split-adjusted), not by today's count — so splits and buybacks don't move the line.
Short interest has its own tab now — open SI →
Source: SEC Form 13F — every institution manager (≥ $100M in 13F securities) reporting a long US-listed position, across ~10,250 filers over 10 years of history. 13F is long-only, US-listed, filed up to 45 days after quarter-end.
| Biggest buyers | Δ shares | Δ value | |
|---|---|---|---|
| VANGUARD PORTFOLIO MANAGEMENT LLC | +9,215,577 | $83M | OPEN |
| VANGUARD CAPITAL MANAGEMENT LLC | +5,382,103 | $48M | OPEN |
| ADAR1 Capital Management, LLC | +2,744,714 | $25M | OPEN |
| Soleus Capital Management, L.P. | +2,002,000 | $18M | OPEN |
| Point72 Asset Management, L.P. | +1,756,516 | $16M | ADD |
| Biggest sellers | Δ shares | Δ value | |
|---|---|---|---|
| VANGUARD GROUP INC | −12,548,944 | $112M | CLOSE |
| ORBIMED ADVISORS LLC | −2,380,715 | $21M | CLOSE |
| SB INVESTMENT ADVISERS (UK) LTD | −2,168,884 | $19M | TRIM |
| Aberdeen Group plc | −1,390,577 | $12M | CLOSE |
| Balyasny Asset Management L.P. | −1,368,474 | $12M | CLOSE |
The 2025-Q4 register is as-filed in ITS OWN share basis; every count below was rebased into the 2026-Q1 basis (no split in between, factor 1). Dollar figures value the SHARE delta at one price — the 2026-03-31 close — rather than differencing two filings' own reported values, which would mix the quarter's price move into the trade figure.
13Fs disclose LONG US-listed positions only — shorts, most derivatives, bonds and non-US listings are invisible here, and a filer's quarter-over-quarter change is not necessarily a trade (index rebalances, in-kind transfers and entity successions all read as flow).
| Quarter | Holders | Shares held | % shares out | Value held |
|---|---|---|---|---|
| 2026-Q1 | 251 | 140,103,078 | 100.4% | $1.3B |
| 2025-Q4 | 218 | 115,839,169 | 83.3% | $698M |
| 2025-Q3 | 213 | 107,461,551 | 77.4% | $614M |
| 2025-Q2 | 213 | 111,483,649 | 80.6% | $562M |
| 2025-Q1 | 234 | 93,282,042 | 68.0% | $605M |
| 2024-Q4 | 208 | 88,802,485 | 64.5% | $652M |
| 2024-Q3 | 197 | 88,690,829 | 64.9% | $664M |
| 2024-Q2 | 215 | 89,253,069 | 65.6% | $794M |
| 2024-Q1 | 210 | 94,483,597 | 70.0% | $958M |
| 2023-Q4 | 206 | 90,535,931 | 67.3% | $911M |
| 2023-Q3 | 214 | 88,504,600 | 65.9% | $829M |
| 2023-Q2 | 227 | 90,799,227 | 67.8% | $2.2B |
| 2023-Q1 | 233 | 88,692,507 | 66.4% | $2.1B |
| 2022-Q4 | 225 | 86,084,146 | 64.7% | $2.2B |
| 2022-Q3 | 221 | 86,959,111 | 65.6% | $1.7B |
| 2022-Q2 | 213 | 85,843,502 | 64.8% | $2.2B |
| 2022-Q1 | 204 | 81,153,082 | 61.3% | $2.1B |
| 2021-Q4 | 210 | 78,286,783 | 59.8% | $3.3B |
| 2021-Q3 | 175 | 74,470,529 | 57.0% | $3.2B |
| 2021-Q2 | 166 | 76,951,120 | 59.2% | $3.6B |
| 2021-Q1 | 157 | 75,667,922 | 59.2% | $3.9B |
| 2020-Q4 | 143 | 78,996,894 | 62.0% | $2.1B |
| 2020-Q3 | 119 | 70,051,007 | 55.1% | $2.4B |
| 2020-Q2 | 112 | 53,528,807 | 45.4% | $2.2B |
| 2020-Q1 | 61 | 37,869,031 | 34.5% | $1.3B |
| 2019-Q4 | 38 | 36,619,648 | 33.4% | $415M |
| VANGUARD GROUP INC | 0% | QII | 2026-03-13 | SEC → |
Source: SEC Schedule 13D/13G structured XML. 13D = active intent; 13G filers report passively (Rule 13d-1(b) QII / 13d-1(c) Passive / 13d-1(d) Exempt) and are not activists. Percent and delta are the group aggregate the filing reports. Structured 13D/G coverage begins December 2024 (SEC XML mandate).
