Institutional 13F holdings sum to 108.5% of shares outstanding. Readings above 100% reflect securities-lending double counting: shares on loan to short sellers are reported long by both the lending institution and the one that bought them. FINRA short interest of 8,896,472 shares (settlement 2026-04-30) accounts for the excess.
| # | Holder | Shares | Value | % of shares out | Δ QoQ |
|---|---|---|---|---|---|
| 1 | RA CAPITAL MANAGEMENT, L.P. | 7,255,9937.3M | $144M | 11.04% | ADD 4% |
| 2 | T. Rowe Price Investment Management, Inc. | 6,605,7806.6M | $131M | 10.05% | ADD 94% |
| 3 | Siren, L.L.C. | 5,802,9705.8M | $115M | 8.83% | ADD 11% |
| 4 | Vestal Point Capital, LP | 5,425,0005.4M | $108M | 8.26% | ADD 52% |
| 5 | Deep Track Capital, LP | 4,062,5004.1M | $81M | 6.18% | ADD 55% |
| 6 | JANUS HENDERSON GROUP PLC | 3,339,2183.3M | $66M | 5.08% | ADD 26% |
| 7 | RED TREE MANAGEMENT, LLC | 3,170,5093.2M | $63M | 4.83% | — |
| 8 | TPG GP A, LLC | 3,010,4253.0M | $60M | 4.58% | — |
| 9 | BlackRock, Inc. | 2,843,7072.8M | $57M | 4.33% | ADD 12% |
| 10 | Omega Fund Management, LLC | 2,392,1502.4M | $48M | 3.64% | ADD 9% |
| 11 | Royalty Pharma Sub-Manager, LLC | 2,388,1502.4M | $47M | 3.63% | ADD 89% |
| 12 | FMR LLC | 2,100,9292.1M | $42M | 3.20% | TRIM -37% |
| 13 | VANGUARD CAPITAL MANAGEMENT LLC | 1,774,2201.8M | $35M | 2.70% | NEW† |
| 14 | CITADEL ADVISORS LLC | 1,659,3421.7M | $33M | 2.53% | ADD 115% |
| 15 | FRANKLIN RESOURCES INC | 1,621,6431.6M | $32M | 2.47% | — |
| 16 | STATE STREET CORP | 1,419,3881.4M | $28M | 2.16% | TRIM -14% |
| 17 | PRICE T ROWE ASSOCIATES INC /MD/ | 1,376,1981.4M | $27M | 2.09% | TRIM -12% |
| 18 | Jones Hill Capital LP | 1,260,1361.3M | $25M | 1.92% | NEW |
| 19 | GEODE CAPITAL MANAGEMENT, LLC | 830,906830,906 | $17M | 1.26% | ADD 2% |
| 20 | FIL Ltd | 679,904679,904 | $14M | 1.03% | TRIM -10% |
| 21 | ORBIMED ADVISORS LLC | 667,300667,300 | $13M | 1.02% | ADD 219% |
| 22 | ACORN CAPITAL ADVISORS, LLC | 643,382643,382 | $13M | 0.98% | — |
| 23 | Aisling Capital Management LP | 568,919568,919 | $11M | 0.87% | — |
| 24 | GOLDMAN SACHS GROUP INC | 558,964558,964 | $11M | 0.85% | — |
| 25 | Palo Alto Investors LP | 550,408550,408 | $11M | 0.84% | ADD 84% |
| 26 | Rock Springs Capital Management LP | 489,095489,095 | $10M | 0.74% | ADD 14% |
| 27 | NAN FUNG TRINITY (HK) LTD | 427,885427,885 | $9M | 0.65% | ADD 15% |
| 28 | DAFNA Capital Management LLC | 421,733421,733 | $8M | 0.64% | ADD 61% |
| 29 | MORGAN STANLEY | 421,219421,219 | $8M | 0.64% | ADD 285% |
| 30 | Ensign Peak Advisors, Inc | 401,444401,444 | $8M | 0.61% | — |
| 31 | ALGERT GLOBAL LLC | 330,833330,833 | $7M | 0.50% | ADD 4% |
| 32 | BANK OF AMERICA CORP /DE/ | 327,989327,989 | $7M | 0.50% | TRIM -2% |
| 33 | VANGUARD PORTFOLIO MANAGEMENT LLC | 290,939290,939 | $6M | 0.44% | NEW |
| 34 | Clearbridge Investments, LLC | 289,221289,221 | $6M | 0.44% | TRIM -3% |
| 35 | D. E. Shaw & Co., Inc. | 276,943276,943 | $6M | 0.42% | ADD 103% |
| 36 | TD Asset Management Inc | 269,718269,718 | $5M | 0.41% | ADD 10% |
| 37 | CHARLES SCHWAB INVESTMENT MANAGEMENT INC | 263,985263,985 | $5M | 0.40% | TRIM -3% |
| 38 | NORTHERN TRUST CORP | 255,571255,571 | $5M | 0.39% | ADD 8% |
| 39 | Candriam S.C.A. | 238,822238,822 | $5M | 0.36% | NEW |
| 40 | Hudson Bay Capital Management LP | 233,703233,703 | $5M | 0.36% | ADD 56% |
| 41 | Woodline Partners LP | 232,690232,690 | $5M | 0.35% | — |
| 42 | VANGUARD FIDUCIARY TRUST CO | 216,753216,753 | $4M | 0.33% | NEW |
| 43 | UBS Group AG | 179,491179,491 | $4M | 0.27% | TRIM -7% |
| 44 | JPMORGAN CHASE & CO | 177,408177,408 | $3M | 0.27% | ADD 953% |
| 45 | MILLENNIUM MANAGEMENT LLC | 156,702156,702 | $3M | 0.24% | TRIM -64% |
| 46 | DEUTSCHE BANK AG\ | 131,656131,656 | $3M | 0.20% | ADD 2% |
| 47 | Atle Fund Management AB | 1,169,6341.2M | $2M | 1.78% | ADD 950% |
| 48 | Napean Trading & Investment Co (Singapore) PTE Ltd | 113,780113,780 | $2M | 0.17% | — |
| 49 | DIMENSIONAL FUND ADVISORS LP | 109,261109,261 | $2M | 0.17% | ADD 167% |
| 50 | Walleye Capital LLC | 108,008108,008 | $2M | 0.16% | NEW |
| 51 | Burkehill Global Management, LP | 100,000100,000 | $2M | 0.15% | NEW |
| 52 | MANUFACTURERS LIFE INSURANCE COMPANY, THE | 90,28590,285 | $2M | 0.14% | TRIM -9% |
| 53 | TWO SIGMA INVESTMENTS, LP | 89,88189,881 | $2M | 0.14% | TRIM -9% |
| 54 | Bank of New York Mellon Corp | 83,26483,264 | $2M | 0.13% | TRIM -6% |
| 55 | Shay Capital LLC | 63,50063,500 | $1M | 0.10% | — |
| 56 | SEVEN GRAND MANAGERS, LLC | 62,50062,500 | $1M | 0.10% | NEW |
| 57 | MARTINGALE ASSET MANAGEMENT L P | 57,53857,538 | $1M | 0.09% | NEW |
| 58 | Mariner, LLC | 55,03755,037 | $1M | 0.08% | TRIM -15% |
| 59 | Swiss National Bank | 55,00055,000 | $1M | 0.08% | ADD 70% |
| 60 | SCS Capital Management LLC | 53,43053,430 | $1M | 0.08% | — |
| 61 | ALPS ADVISORS INC | 51,58051,580 | $1M | 0.08% | ADD 15% |
| 62 | AMERICAN CENTURY COMPANIES INC | 51,55351,553 | $1M | 0.08% | ADD 110% |
| 63 | Swiss Life Asset Management Ltd | 8,6518,651 | $978,718 | 0.01% | NEW |
| 64 | Nuveen, LLC | 43,98443,984 | $874,841 | 0.07% | ADD 13% |
| 65 | RHUMBLINE ADVISERS | 40,74740,747 | $810,450 | 0.06% | — |
| 66 | BNP PARIBAS FINANCIAL MARKETS | 40,39640,396 | $803,476 | 0.06% | TRIM -58% |
| 67 | HRT FINANCIAL LP | 40,20940,209 | $799,000 | 0.06% | TRIM -20% |
| 68 | Rafferty Asset Management, LLC | 38,00338,003 | $755,880 | 0.06% | TRIM -36% |
| 69 | Invesco Ltd. | 31,65031,650 | $629,518 | 0.05% | ADD 51% |
| 70 | Bleakley Financial Group, LLC | 29,33129,331 | $583,394 | 0.04% | — |
| 71 | BARCLAYS PLC | 28,82328,823 | $573,289 | 0.04% | TRIM -72% |
| 72 | PRUDENTIAL FINANCIAL INC | 27,33027,330 | $543,594 | 0.04% | NEW |
| 73 | ALLIANCEBERNSTEIN L.P. | 31,83031,830 | $535,699 | 0.05% | ADD 9% |
| 74 | Optimize Financial Inc | 23,99623,996 | $477,280 | 0.04% | NEW |
| 75 | VANGUARD ASSET MANAGEMENT, Ltd | 23,94323,943 | $476,226 | 0.04% | NEW |
| 76 | Caption Management, LLC | 23,00023,000 | $457,470 | 0.04% | — |
| 77 | Legal & General Group Plc | 22,22722,227 | $442,095 | 0.03% | ADD 39% |
| 78 | Creative Planning | 21,81621,816 | $433,920 | 0.03% | — |
| 79 | Balyasny Asset Management L.P. | 21,79721,797 | $433,542 | 0.03% | NEW |
| 80 | Russell Investments Group, Ltd. | 19,89319,893 | $395,670 | 0.03% | TRIM -65% |
| 81 | MetLife Investment Management, LLC | 18,36018,360 | $365,180 | 0.03% | TRIM -15% |
| 82 | VOYA INVESTMENT MANAGEMENT LLC | 18,00818,008 | $358,179 | 0.03% | — |
| 83 | Quantinno Capital Management LP | 17,61217,612 | $350,303 | 0.03% | ADD 16% |
| 84 | MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd. | 17,66617,666 | $349,130 | 0.03% | TRIM -3% |
| 85 | PROFUND ADVISORS LLC | 17,12817,128 | $340,676 | 0.03% | TRIM -23% |
| 86 | CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM | 16,92316,923 | $336,598 | 0.03% | — |
| 87 | AMERIPRISE FINANCIAL INC | 15,81815,818 | $314,620 | 0.02% | — |
| 88 | WELLS FARGO & COMPANY/MN | 14,97714,977 | $297,893 | 0.02% | TRIM -58% |
| 89 | Virtus Investment Advisers, LLC | 14,75514,755 | $293,477 | 0.02% | ADD 3% |
| 90 | Ascent Group, LLC | 14,15214,152 | $281,483 | 0.02% | — |
| 91 | LPL Financial LLC | 14,10614,106 | $280,568 | 0.02% | — |
| 92 | Susquehanna Portfolio Strategies, LLC | 14,01714,017 | $278,798 | 0.02% | NEW |
| 93 | ADAR1 Capital Management, LLC | 13,70513,705 | $272,592 | 0.02% | — |
| 94 | VARCOV Co. | 13,29213,292 | $264,378 | 0.02% | TRIM -15% |
| 95 | AQR CAPITAL MANAGEMENT LLC | 13,18613,186 | $262,269 | 0.02% | NEW |
| 96 | Vanguard Global Advisers, LLC | 12,73612,736 | $253,319 | 0.02% | NEW |
| 97 | China Universal Asset Management Co., Ltd. | 11,24411,244 | $223,643 | 0.02% | ADD 26% |
| 98 | LETKO, BROSSEAU & ASSOCIATES INC | 11,17511,175 | $222,271 | 0.02% | — |
| 99 | SUSQUEHANNA INTERNATIONAL GROUP, LLP | 10,94410,944 | $217,676 | 0.02% | TRIM -64% |
| 100 | Squarepoint Ops LLC | 10,77110,771 | $214,235 | 0.02% | NEW |
† a filer this large appearing as NEW usually reflects a renamed or succeeded filing entity (the old name stops filing the same quarter) — not a brand-new position.
Rows rank by the value each filer reported — 13F filers price their own positions, so a holder with fewer shares can rank just above one with more.
| Holder | 2024-Q4 | 2025-Q1 | 2025-Q2 | 2025-Q3 | 2025-Q4 | 2026-Q1 | Latest value |
|---|---|---|---|---|---|---|---|
| RA CAPITAL MANAGEMENT, L.P. | 7.0M | 7.0M | 7.0M | 7.0M | 7.0M | 7.3M | $144M |
| T. Rowe Price Investment Management, Inc. | — | — | — | 465K | 3.4M | 6.6M | $131M |
| Siren, L.L.C. | 3.3M | 3.3M | 5.2M | 5.2M | 5.2M | 5.8M | $115M |
| Vestal Point Capital, LP | 710K | 2.8M | 5.0M | 5.1M | 3.6M | 5.4M | $108M |
| Deep Track Capital, LP | 2.0M | 2.0M | 2.4M | 2.4M | 2.6M | 4.1M | $81M |
| JANUS HENDERSON GROUP PLC | 864K | 1.3M | — | — | 2.6M | 3.3M | $66M |
| RED TREE MANAGEMENT, LLC | 3.2M | 3.2M | 3.2M | 3.2M | 3.2M | 3.2M | $63M |
| TPG GP A, LLC | 3.0M | 3.0M | 3.0M | 3.0M | 3.0M | 3.0M | $60M |
| BlackRock, Inc. | 1.3M | 1.3M | 2.1M | 2.3M | 2.5M | 2.8M | $57M |
| Omega Fund Management, LLC | 2.2M | 2.2M | 2.2M | 2.2M | 2.2M | 2.4M | $48M |
| Royalty Pharma Sub-Manager, LLC | — | — | — | — | 1.3M | 2.4M | $47M |
| FMR LLC | 2.1M | 3.6M | 2.5M | 2.7M | 3.3M | 2.1M | $42M |
| VANGUARD CAPITAL MANAGEMENT LLC | — | — | — | — | — | 1.8M | $35M |
| CITADEL ADVISORS LLC | — | — | 222K | — | 772K | 1.7M | $33M |
| FRANKLIN RESOURCES INC | — | 14K | — | — | 1.6M | 1.6M | $32M |
Shares reported each quarter by the largest current holders. Green = added vs the prior filing, red = trimmed, grey = roughly held; a dash is a quarter with no 13F on file (a gap is skipped, not read as an exit). 13F is filed up to 45 days after quarter-end. Up to 40 quarters (10 years) available — use the selector above.
Line breaks are honest gaps — quarters with no 13F record, or a suppressed % — never interpolated. % of shares outstanding uses today's share count for every quarter (the only count stored).
| Quarter | Holders | Shares held | % shares out | Value held |
|---|---|---|---|---|
| 2026-Q1 | 147 | 71,261,744 | 108.5% | $1.4B |
| 2025-Q4 | 141 | 58,518,644 | 89.1% | $1.0B |
| 2025-Q3 | 114 | 50,223,485 | 76.4% | $790M |
| 2025-Q2 | 106 | 51,172,768 | 77.9% | $475M |
| 2025-Q1 | 93 | 48,610,442 | 74.0% | $634M |
| 2024-Q4 | 84 | 46,709,070 | 71.1% | $814M |
| 2024-Q3 | 69 | 36,418,294 | 55.4% | $928M |
Short interest has its own tab now — open SI →
Source: SEC Form 13F — every institution manager (≥ $100M in 13F securities) reporting a long US-listed position, across ~10,250 filers over 10 years of history. 13F is long-only, US-listed, filed up to 45 days after quarter-end.
| Deep Track Capital, LP | 6.21% | Passive | 2026-03-31 | SEC → |
| BlackRock, Inc. | 5.4% | QII | 2026-06-30 | SEC → |
Source: SEC Schedule 13D/13G structured XML. 13D = active intent; 13G filers report passively (Rule 13d-1(b) QII / 13d-1(c) Passive / 13d-1(d) Exempt) and are not activists. Percent and delta are the group aggregate the filing reports. Structured 13D/G coverage begins December 2024 (SEC XML mandate).
