| # | Holder | Shares | Value | % of shares out | Δ QoQ |
|---|---|---|---|---|---|
| 1 | L1 Capital Pty Ltd | 7,789,0007.8M | $43M | 8.00% | ADD 177% |
| 2 | Nantahala Capital Management, LLC | 5,530,9105.5M | $31M | 5.68% | ADD 119% |
| 3 | Sio Capital Management, LLC | 5,382,9655.4M | $30M | 5.53% | ADD 187% |
| 4 | BlackRock, Inc. | 4,889,1534.9M | $27M | 5.02% | ADD 141% |
| 5 | ADAR1 Capital Management, LLC | 3,789,1323.8M | $21M | 3.89% | ADD 206% |
| 6 | PERCEPTIVE ADVISORS LLC | 3,425,4763.4M | $19M | 3.52% | ADD 104% |
| 7 | BVF INC/IL | 3,000,0003.0M | $17M | 3.08% | NEW† |
| 8 | RA CAPITAL MANAGEMENT, L.P. | 3,000,0003.0M | $17M | 3.08% | NEW† |
| 9 | FMR LLC | 2,901,3502.9M | $16M | 2.98% | ADD 7221% |
| 10 | Ensign Peak Advisors, Inc | 2,087,1112.1M | $12M | 2.14% | ADD 49% |
| 11 | GEODE CAPITAL MANAGEMENT, LLC | 1,841,2301.8M | $10M | 1.89% | ADD 131% |
| 12 | Rhenman & Partners Asset Management AB | 1,438,6251.4M | $8M | 1.48% | ADD 16% |
| 13 | UBS Group AG | 1,406,8361.4M | $8M | 1.45% | ADD 173% |
| 14 | STATE STREET CORP | 1,223,1071.2M | $7M | 1.26% | ADD 242% |
| 15 | Vestal Point Capital, LP | 1,144,7801.1M | $6M | 1.18% | NEW |
| 16 | Affinity Asset Advisors, LLC | 912,397912,397 | $5M | 0.94% | ADD 87% |
| 17 | Oasis Management Co Ltd. | 865,565865,565 | $5M | 0.89% | ADD 1040% |
| 18 | VANGUARD CAPITAL MANAGEMENT LLC | 857,803857,803 | $5M | 0.88% | NEW |
| 19 | SUVRETTA CAPITAL MANAGEMENT, LLC | 850,000850,000 | $5M | 0.87% | NEW |
| 20 | Alyeska Investment Group, L.P. | 750,000750,000 | $4M | 0.77% | NEW |
| 21 | Velan Capital Investment Management LP | 700,000700,000 | $4M | 0.72% | NEW |
| 22 | AMERIPRISE FINANCIAL INC | 653,330653,330 | $4M | 0.67% | NEW |
| 23 | Militia Capital Management LLC | 605,000605,000 | $3M | 0.62% | NEW |
| 24 | NORTHERN TRUST CORP | 591,046591,046 | $3M | 0.61% | ADD 132% |
| 25 | Capricorn Fund Managers Ltd | 500,525500,525 | $3M | 0.51% | NEW |
| 26 | VANGUARD FIDUCIARY TRUST CO | 476,344476,344 | $3M | 0.49% | NEW |
| 27 | VANGUARD PORTFOLIO MANAGEMENT LLC | 469,583469,583 | $3M | 0.48% | NEW |
| 28 | LMR Partners LLP | 450,000450,000 | $2M | 0.46% | NEW |
| 29 | GOLDMAN SACHS GROUP INC | 415,493415,493 | $2M | 0.43% | ADD 172% |
| 30 | Anson Funds Management LP | 405,652405,652 | $2M | 0.42% | NEW |
| 31 | Eversept Partners, LP | 401,187401,187 | $2M | 0.41% | TRIM -20% |
| 32 | Woodline Partners LP | 400,000400,000 | $2M | 0.41% | NEW |
| 33 | Ikarian Capital, LLC | 366,571366,571 | $2M | 0.38% | NEW |
| 34 | SEVEN GRAND MANAGERS, LLC | 325,000325,000 | $2M | 0.33% | NEW |
| 35 | Avidity Partners Management LP | 300,000300,000 | $2M | 0.31% | NEW |
| 36 | Nuveen, LLC | 307,864307,864 | $2M | 0.32% | ADD 621% |
| 37 | Jain Global LLC | 303,286303,286 | $2M | 0.31% | NEW |
| 38 | SummitTX Capital, L.P. | 238,947238,947 | $1M | 0.25% | NEW |
| 39 | CHARLES SCHWAB INVESTMENT MANAGEMENT INC | 227,910227,910 | $1M | 0.23% | ADD 126% |
| 40 | MILLENNIUM MANAGEMENT LLC | 196,221196,221 | $1M | 0.20% | ADD 129% |
| 41 | Symmetry Investments LP | 182,837182,837 | $1M | 0.19% | ADD 15% |
| 42 | Bank of New York Mellon Corp | 170,653170,653 | $947,125 | 0.18% | ADD 126% |
| 43 | MORGAN STANLEY | 165,358165,358 | $917,736 | 0.17% | ADD 106% |
| 44 | Ghisallo Capital Management LLC | 150,000150,000 | $832,500 | 0.15% | NEW |
| 45 | BOOTHBAY FUND MANAGEMENT, LLC | 137,510137,510 | $763,181 | 0.14% | NEW |
| 46 | SUSQUEHANNA INTERNATIONAL GROUP, LLP | 123,580123,580 | $685,869 | 0.13% | NEW |
| 47 | AWM Investment Company, Inc. | 115,000115,000 | $638,250 | 0.12% | NEW |
| 48 | BARCLAYS PLC | 109,922109,922 | $610,067 | 0.11% | TRIM -26% |
| 49 | BEARING POINT CAPITAL, LLC | 101,560101,560 | $563,658 | 0.10% | ADD 6% |
| 50 | Empower Advisory Group, LLC | 101,082101,082 | $561,005 | 0.10% | NEW |
| 51 | Marex Group plc | 94,93494,934 | $526,884 | 0.10% | NEW |
| 52 | Persistent Asset Partners Ltd | 85,91385,913 | $476,817 | 0.09% | ADD 268% |
| 53 | MACQUARIE GROUP LTD | 81,24681,246 | $450,915 | 0.08% | ADD 141% |
| 54 | Quinn Opportunity Partners LLC | 75,00075,000 | $416,250 | 0.08% | NEW |
| 55 | DEUTSCHE BANK AG\ | 71,92271,922 | $399,167 | 0.07% | ADD 149% |
| 56 | AQR CAPITAL MANAGEMENT LLC | 70,77470,774 | $392,794 | 0.07% | ADD 222% |
| 57 | BANK OF AMERICA CORP /DE/ | 68,75568,755 | $381,590 | 0.07% | ADD 13% |
| 58 | Susquehanna Portfolio Strategies, LLC | 66,32566,325 | $368,104 | 0.07% | NEW |
| 59 | BRIDGEWAY CAPITAL MANAGEMENT, LLC | 60,00060,000 | $333,000 | 0.06% | ADD 71% |
| 60 | RHUMBLINE ADVISERS | 59,18359,183 | $328,459 | 0.06% | ADD 57% |
| 61 | PRICE T ROWE ASSOCIATES INC /MD/ | 56,23556,235 | $313,000 | 0.06% | ADD 146% |
| 62 | Dynamic Technology Lab Private Ltd | 51,05851,058 | $283,372 | 0.05% | NEW |
| 63 | Shay Capital LLC | 50,91350,913 | $282,567 | 0.05% | NEW |
| 64 | Covalis Capital LLP | 50,00050,000 | $277,500 | 0.05% | NEW |
| 65 | WELLS FARGO & COMPANY/MN | 49,72849,728 | $275,990 | 0.05% | ADD 34% |
| 66 | FIFTH LANE CAPITAL, LP | 47,50047,500 | $263,625 | 0.05% | NEW |
| 67 | MetLife Investment Management, LLC | 44,53644,536 | $247,175 | 0.05% | ADD 571% |
| 68 | Invesco Ltd. | 40,52240,522 | $224,897 | 0.04% | ADD 11% |
| 69 | One68 Global Capital, LLC | 39,78039,780 | $220,779 | 0.04% | NEW |
| 70 | MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd. | 37,19137,191 | $206,410 | 0.04% | ADD 135% |
| 71 | JPMORGAN CHASE & CO | 32,41332,413 | $165,631 | 0.03% | TRIM -98% |
| 72 | CITADEL ADVISORS LLC | 28,82128,821 | $159,957 | 0.03% | NEW |
| 73 | Vanguard Global Advisers, LLC | 28,14528,145 | $156,205 | 0.03% | NEW |
| 74 | Wellington Shields & Co., LLC | 27,28427,284 | $151,426 | 0.03% | — |
| 75 | Virtu Financial LLC | 26,57626,576 | $147,000 | 0.03% | NEW |
| 76 | GDS Wealth Management | 26,10026,100 | $144,855 | 0.03% | NEW |
| 77 | Quadrature Capital Ltd | 23,59023,590 | $130,925 | 0.02% | NEW |
| 78 | Police & Firemen's Retirement System of New Jersey | 21,26321,263 | $118,010 | 0.02% | ADD 180% |
| 79 | XTX Topco Ltd | 20,73220,732 | $115,063 | 0.02% | NEW |
| 80 | MANUFACTURERS LIFE INSURANCE COMPANY, THE | 20,06320,063 | $111,350 | 0.02% | NEW |
| 81 | LPL Financial LLC | 17,84517,845 | $99,040 | 0.02% | ADD 3% |
| 82 | BNP PARIBAS FINANCIAL MARKETS | 17,63717,637 | $97,885 | 0.02% | ADD 112% |
| 83 | NEW YORK STATE COMMON RETIREMENT FUND | 17,40017,400 | $96,570 | 0.02% | ADD 255% |
| 84 | TWO SIGMA INVESTMENTS, LP | 17,33517,335 | $96,209 | 0.02% | NEW |
| 85 | Numerai GP LLC | 16,75316,753 | $92,979 | 0.02% | NEW |
| 86 | Cetera Investment Advisers | 16,55016,550 | $91,853 | 0.02% | ADD 65% |
| 87 | INTECH INVESTMENT MANAGEMENT LLC | 14,37914,379 | $79,803 | 0.01% | ADD 7% |
| 88 | BARTLETT & CO. WEALTH MANAGEMENT LLC | 12,90012,900 | $79,077 | 0.01% | ADD 174% |
| 89 | VOYA INVESTMENT MANAGEMENT LLC | 13,30413,304 | $73,837 | 0.01% | NEW |
| 90 | PRUDENTIAL FINANCIAL INC | 13,18513,185 | $73,177 | 0.01% | NEW |
| 91 | ProShare Advisors LLC | 11,50111,501 | $63,830 | 0.01% | NEW |
| 92 | Abel Hall, LLC | 11,34511,345 | $62,965 | 0.01% | NEW |
| 93 | Duncan Williams Asset Management, LLC | 10,87010,870 | $60,329 | 0.01% | — |
| 94 | CAPTRUST FINANCIAL ADVISORS | 10,42710,427 | $57,858 | 0.01% | NEW |
| 95 | Russell Investments Group, Ltd. | 9,7609,760 | $54,168 | 0.01% | ADD 49% |
| 96 | CITIGROUP INC | 7,3787,378 | $40,948 | 0.01% | TRIM -96% |
| 97 | Legal & General Group Plc | 6,9436,943 | $38,534 | 0.01% | ADD 101% |
| 98 | Tower Research Capital LLC (TRC) | 6,5586,558 | $36,397 | 0.01% | ADD 167% |
| 99 | Ameritas Investment Partners, Inc. | 6,4916,491 | $36,025 | 0.01% | ADD 327% |
| 100 | FNY Investment Advisers, LLC | 5,0005,000 | $27,000 | 0.01% | NEW |
† a filer this large appearing as NEW usually reflects a renamed or succeeded filing entity (the old name stops filing the same quarter) — not a brand-new position.
Rows rank by the value each filer reported — 13F filers price their own positions, so a holder with fewer shares can rank just above one with more.
| Holder | 2024-Q4 | 2025-Q1 | 2025-Q2 | 2025-Q3 | 2025-Q4 | 2026-Q1 | Latest value |
|---|---|---|---|---|---|---|---|
| L1 Capital Pty Ltd | 2.8M | 6.9M | 2.8M | 6.8M | 2.8M | 7.8M | $43M |
| Nantahala Capital Management, LLC | 1.7M | 1.7M | 1.7M | 1.9M | 2.5M | 5.5M | $31M |
| Sio Capital Management, LLC | 2.4M | 2.4M | 1.8M | 1.8M | 1.9M | 5.4M | $30M |
| BlackRock, Inc. | — | 395K | 1.0M | 1.9M | 2.0M | 4.9M | $27M |
| ADAR1 Capital Management, LLC | 1.1M | 1.7M | 532K | 198K | 1.2M | 3.8M | $21M |
| PERCEPTIVE ADVISORS LLC | 2.1M | 2.1M | 1.7M | 1.7M | 1.7M | 3.4M | $19M |
| BVF INC/IL | — | — | — | — | — | 3.0M | $17M |
| RA CAPITAL MANAGEMENT, L.P. | — | — | — | — | — | 3.0M | $17M |
| FMR LLC | 584K | 2K | 33K | 37K | 40K | 2.9M | $16M |
| Ensign Peak Advisors, Inc | 1.0M | 1.4M | 1.4M | 1.4M | 1.4M | 2.1M | $12M |
| GEODE CAPITAL MANAGEMENT, LLC | — | 165K | 420K | 749K | 796K | 1.8M | $10M |
| Rhenman & Partners Asset Management AB | — | 1.3M | 1.3M | 1.3M | 1.2M | 1.4M | $8M |
| UBS Group AG | — | 130K | 301K | 221K | 516K | 1.4M | $8M |
| STATE STREET CORP | — | 89K | 189K | 313K | 358K | 1.2M | $7M |
| Vestal Point Capital, LP | — | — | — | — | — | 1.1M | $6M |
Shares reported each quarter by the largest current holders. Green = added vs the prior filing, red = trimmed, grey = roughly held; a dash is a quarter with no 13F on file (a gap is skipped, not read as an exit). 13F is filed up to 45 days after quarter-end. Up to 40 quarters (10 years) available — use the selector above.
Line breaks are honest gaps — quarters with no 13F record, or a suppressed % — never interpolated. % of shares outstanding uses today's share count for every quarter (the only count stored).
| Quarter | Holders | Shares held | % shares out | Value held |
|---|---|---|---|---|
| 2026-Q1 | 127 | 64,506,279 | 66.3% | $358M |
| 2025-Q4 | 74 | 21,817,667 | 22.4% | $109M |
| 2025-Q3 | 66 | 25,126,131 | 25.8% | $110M |
| 2025-Q2 | 60 | 17,136,184 | 17.6% | $80M |
| 2025-Q1 | 55 | 22,060,522 | 22.7% | $80M |
| 2024-Q4 | 27 | 16,269,470 | 16.7% | $91M |
Short interest has its own tab now — open SI →
Source: SEC Form 13F — every institution manager (≥ $100M in 13F securities) reporting a long US-listed position, across ~10,250 filers over 10 years of history. 13F is long-only, US-listed, filed up to 45 days after quarter-end.
