Institutional 13F holdings sum to 105.7% of shares outstanding. Readings above 100% reflect securities-lending double counting: shares on loan to short sellers are reported long by both the lending institution and the one that bought them. FINRA short interest of 10,723,872 shares (settlement 2026-04-30) accounts for the excess.
| # | Holder | Shares | Value | % of shares out | Δ QoQ |
|---|---|---|---|---|---|
| 1 | RA CAPITAL MANAGEMENT, L.P. | 4,178,8234.2M | $38M | 13.33% | — |
| 2 | Alpha Wave Global, LP | 3,612,2113.6M | $33M | 11.52% | — |
| 3 | FMR LLC | 3,597,7843.6M | $33M | 11.47% | TRIM -3% |
| 4 | Versant Venture Management, LLC | 2,656,8882.7M | $24M | 8.47% | — |
| 5 | BlackRock, Inc. | 1,976,8232.0M | $18M | 6.30% | ADD 17% |
| 6 | Squarepoint Ops LLC | 1,506,4991.5M | $14M | 4.80% | ADD 206% |
| 7 | Aberdeen Group plc | 1,156,1611.2M | $11M | 3.69% | ADD 136% |
| 8 | FRANKLIN RESOURCES INC | 1,100,1481.1M | $10M | 3.51% | — |
| 9 | VANGUARD CAPITAL MANAGEMENT LLC | 1,041,1881.0M | $10M | 3.32% | NEW† |
| 10 | 72 Investment Holdings, LLC | 1,017,7511.0M | $9M | 3.25% | — |
| 11 | MILLENNIUM MANAGEMENT LLC | 869,257869,257 | $8M | 2.77% | ADD 61% |
| 12 | C WorldWide Group Holding A/S | 700,000700,000 | $6M | 2.23% | ADD 40% |
| 13 | MORGAN STANLEY | 648,848648,848 | $6M | 2.07% | ADD 178% |
| 14 | BANK OF AMERICA CORP /DE/ | 547,534547,534 | $5M | 1.75% | ADD 377% |
| 15 | GEODE CAPITAL MANAGEMENT, LLC | 524,574524,574 | $5M | 1.67% | ADD 14% |
| 16 | Blue Owl Capital Holdings LP | 506,722506,722 | $5M | 1.62% | ADD 238% |
| 17 | TWO SIGMA INVESTMENTS, LP | 500,997500,997 | $5M | 1.60% | ADD 71% |
| 18 | PRIMECAP MANAGEMENT CO/CA/ | 472,200472,200 | $4M | 1.51% | ADD 27% |
| 19 | Alyeska Investment Group, L.P. | 463,495463,495 | $4M | 1.48% | — |
| 20 | TANG CAPITAL MANAGEMENT LLC | 451,114451,114 | $4M | 1.44% | TRIM -48% |
| 21 | STATE STREET CORP | 402,225402,225 | $4M | 1.28% | TRIM -21% |
| 22 | SILVERCREST ASSET MANAGEMENT GROUP LLC | 283,206283,206 | $3M | 0.90% | ADD 62% |
| 23 | Rock Springs Capital Management LP | 257,460257,460 | $2M | 0.82% | — |
| 24 | DISCOVERY CAPITAL MANAGEMENT, LLC / CT | 241,300241,300 | $2M | 0.77% | — |
| 25 | XTX Topco Ltd | 221,037221,037 | $2M | 0.70% | ADD 158% |
| 26 | Trexquant Investment LP | 214,408214,408 | $2M | 0.68% | ADD 24% |
| 27 | NORTHERN TRUST CORP | 185,624185,624 | $2M | 0.59% | ADD 15% |
| 28 | GOLDMAN SACHS GROUP INC | 183,756183,756 | $2M | 0.59% | TRIM -29% |
| 29 | MARSHALL WACE, LLP | 160,264160,264 | $1M | 0.51% | ADD 355% |
| 30 | Y-Intercept (Hong Kong) Ltd | 158,715158,715 | $1M | 0.51% | ADD 291% |
| 31 | UBS Group AG | 156,989156,989 | $1M | 0.50% | TRIM -90% |
| 32 | VANGUARD PORTFOLIO MANAGEMENT LLC | 153,004153,004 | $1M | 0.49% | NEW |
| 33 | BARCLAYS PLC | 151,387151,387 | $1M | 0.48% | ADD 109% |
| 34 | VANGUARD FIDUCIARY TRUST CO | 149,987149,987 | $1M | 0.48% | NEW |
| 35 | CHARLES SCHWAB INVESTMENT MANAGEMENT INC | 149,544149,544 | $1M | 0.48% | TRIM -12% |
| 36 | HRT FINANCIAL LP | 145,506145,506 | $1M | 0.46% | TRIM -15% |
| 37 | AQR CAPITAL MANAGEMENT LLC | 140,305140,305 | $1M | 0.45% | NEW |
| 38 | SCOGGIN MANAGEMENT LP | 125,000125,000 | $1M | 0.40% | — |
| 39 | JACOBS LEVY EQUITY MANAGEMENT, INC | 117,011117,011 | $1M | 0.37% | TRIM -57% |
| 40 | SUSQUEHANNA INTERNATIONAL GROUP, LLP | 110,304110,304 | $1M | 0.35% | ADD 63% |
| 41 | Jump Financial, LLC | 100,200100,200 | $916,830 | 0.32% | ADD 145% |
| 42 | Affinity Asset Advisors, LLC | 100,000100,000 | $915,000 | 0.32% | NEW |
| 43 | Woodline Partners LP | 100,000100,000 | $915,000 | 0.32% | — |
| 44 | ROYAL BANK OF CANADA | 95,23595,235 | $870,000 | 0.30% | ADD 121% |
| 45 | KBC Group NV | 90,42590,425 | $827,000 | 0.29% | NEW |
| 46 | Rockefeller Capital Management L.P. | 73,34073,340 | $671,062 | 0.23% | ADD 53% |
| 47 | Vivaldi Capital Management LP | 64,05264,052 | $586,076 | 0.20% | — |
| 48 | ExodusPoint Capital Management, LP | 57,57757,577 | $526,830 | 0.18% | ADD 149% |
| 49 | AXQ CAPITAL, LP | 54,56854,568 | $499,297 | 0.17% | ADD 175% |
| 50 | Bank of New York Mellon Corp | 49,79749,797 | $455,638 | 0.16% | ADD 13% |
| 51 | Quantinno Capital Management LP | 47,64847,648 | $435,979 | 0.15% | NEW |
| 52 | Virtus Investment Advisers, LLC | 46,54246,542 | $425,859 | 0.15% | ADD 24% |
| 53 | DIMENSIONAL FUND ADVISORS LP | 41,87441,874 | $381,339 | 0.13% | TRIM -63% |
| 54 | Nuveen, LLC | 40,19040,190 | $367,739 | 0.13% | — |
| 55 | PRICE T ROWE ASSOCIATES INC /MD/ | 39,62139,621 | $364,000 | 0.13% | TRIM -96% |
| 56 | Schonfeld Strategic Advisors LLC | 39,28539,285 | $359,456 | 0.13% | NEW |
| 57 | Virtu Financial LLC | 38,52038,520 | $352,000 | 0.12% | ADD 212% |
| 58 | Connor, Clark & Lunn Investment Management Ltd. | 37,31437,314 | $341,423 | 0.12% | NEW |
| 59 | Corient Private Wealth LLC | 37,25937,259 | $340,922 | 0.12% | NEW |
| 60 | Jain Global LLC | 37,19237,192 | $340,307 | 0.12% | NEW |
| 61 | Knott David M Jr | 33,73233,732 | $308,648 | 0.11% | TRIM -37% |
| 62 | LOS ANGELES CAPITAL MANAGEMENT LLC | 33,07133,071 | $302,601 | 0.11% | ADD 751% |
| 63 | Engineers Gate Manager LP | 32,24332,243 | $295,023 | 0.10% | ADD 4% |
| 64 | Voleon Capital Management LP | 29,99229,992 | $274,427 | 0.10% | NEW |
| 65 | KESTRA PRIVATE WEALTH SERVICES, LLC | 27,30427,304 | $249,832 | 0.09% | — |
| 66 | UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC | 25,78825,788 | $235,960 | 0.08% | ADD 37% |
| 67 | TD Asset Management Inc | 24,61024,610 | $225,182 | 0.08% | TRIM -9% |
| 68 | State of Alaska, Department of Revenue | 24,15924,159 | $221,000 | 0.08% | ADD 26% |
| 69 | ALLIANCEBERNSTEIN L.P. | 13,38613,386 | $214,176 | 0.04% | — |
| 70 | RHUMBLINE ADVISERS | 22,47622,476 | $205,647 | 0.07% | ADD 11% |
| 71 | HSBC HOLDINGS PLC | 21,89421,894 | $203,833 | 0.07% | NEW |
| 72 | Swiss National Bank | 21,80021,800 | $199,470 | 0.07% | — |
| 73 | DNB Asset Management AS | 20,78020,780 | $190,137 | 0.07% | ADD 8% |
| 74 | ACADIAN ASSET MANAGEMENT LLC | 20,61120,611 | $188,000 | 0.07% | NEW |
| 75 | IEQ CAPITAL, LLC | 18,80718,807 | $172,084 | 0.06% | NEW |
| 76 | KELLEHER FINANCIAL ADVISORS | 17,02317,023 | $155,760 | 0.05% | — |
| 77 | STATE OF WISCONSIN INVESTMENT BOARD | 16,22016,220 | $148,413 | 0.05% | TRIM -50% |
| 78 | PRELUDE CAPITAL MANAGEMENT, LLC | 16,07216,072 | $147,059 | 0.05% | TRIM -91% |
| 79 | Marex Group plc | 16,00016,000 | $146,400 | 0.05% | — |
| 80 | DEUTSCHE BANK AG\ | 15,28015,280 | $139,812 | 0.05% | ADD 11% |
| 81 | Rinkey Investments | 15,18615,186 | $138,952 | 0.05% | — |
| 82 | RAYMOND JAMES FINANCIAL INC | 15,00815,008 | $137,324 | 0.05% | ADD 6% |
| 83 | Invesco Ltd. | 14,92714,927 | $136,582 | 0.05% | ADD 6% |
| 84 | WELLS FARGO & COMPANY/MN | 13,97213,972 | $127,844 | 0.04% | TRIM -39% |
| 85 | HARBOR CAPITAL ADVISORS, INC. | 13,90913,909 | $127,000 | 0.04% | TRIM -42% |
| 86 | Brevan Howard Capital Management LP | 13,57013,570 | $124,166 | 0.04% | TRIM -12% |
| 87 | CWM, LLC | 12,24012,240 | $111,996 | 0.04% | ADD 17139% |
| 88 | Russell Investments Group, Ltd. | 10,93410,934 | $100,051 | 0.03% | TRIM -29% |
| 89 | Janney Montgomery Scott LLC | 10,81410,814 | $99,000 | 0.03% | NEW |
| 90 | CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM | 10,81810,818 | $98,985 | 0.03% | TRIM -10% |
| 91 | AlphaCore Capital LLC | 10,00310,003 | $91,527 | 0.03% | — |
| 92 | JPMORGAN CHASE & CO | 10,88310,883 | $90,220 | 0.03% | ADD 15% |
| 93 | MetLife Investment Management, LLC | 9,7879,787 | $89,551 | 0.03% | — |
| 94 | MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd. | 9,3719,371 | $85,745 | 0.03% | ADD 24% |
| 95 | Vanguard Global Advisers, LLC | 8,6368,636 | $79,019 | 0.03% | NEW |
| 96 | Legal & General Group Plc | 8,1698,169 | $74,746 | 0.03% | TRIM -24% |
| 97 | SIMPLEX TRADING, LLC | 6,9276,927 | $63,382 | 0.02% | NEW |
| 98 | VANGUARD ASSET MANAGEMENT, Ltd | 6,1036,103 | $55,842 | 0.02% | NEW |
| 99 | Zurcher Kantonalbank (Zurich Cantonalbank) | 6,0996,099 | $55,806 | 0.02% | ADD 2184% |
| 100 | NEW YORK STATE COMMON RETIREMENT FUND | 5,7935,793 | $53,006 | 0.02% | TRIM -13% |
† a filer this large appearing as NEW usually reflects a renamed or succeeded filing entity (the old name stops filing the same quarter) — not a brand-new position.
Rows rank by the value each filer reported — 13F filers price their own positions, so a holder with fewer shares can rank just above one with more.
| Holder | 2024-Q4 | 2025-Q1 | 2025-Q2 | 2025-Q3 | 2025-Q4 | 2026-Q1 | Latest value |
|---|---|---|---|---|---|---|---|
| RA CAPITAL MANAGEMENT, L.P. | 4.2M | 4.2M | 4.2M | 4.2M | 4.2M | 4.2M | $38M |
| Alpha Wave Global, LP | 3.6M | 3.6M | 3.6M | 3.6M | 3.6M | 3.6M | $33M |
| FMR LLC | 212 | 81K | 82K | 87K | 3.7M | 3.6M | $33M |
| Versant Venture Management, LLC | — | — | — | 2.7M | 2.7M | 2.7M | $24M |
| BlackRock, Inc. | 1.2M | 1.2M | 1.4M | 1.5M | 1.7M | 2.0M | $18M |
| Squarepoint Ops LLC | — | 28K | 10K | 90K | 492K | 1.5M | $14M |
| Aberdeen Group plc | — | — | 74K | 131K | 489K | 1.2M | $11M |
| FRANKLIN RESOURCES INC | — | 9K | 9K | 587K | 1.1M | 1.1M | $10M |
| VANGUARD CAPITAL MANAGEMENT LLC | — | — | — | — | — | 1.0M | $10M |
| 72 Investment Holdings, LLC | 1.0M | 1.0M | 1.0M | 1.0M | 1.0M | 1.0M | $9M |
| MILLENNIUM MANAGEMENT LLC | — | — | — | — | 539K | 869K | $8M |
| C WorldWide Group Holding A/S | — | — | — | — | 500K | 700K | $6M |
| MORGAN STANLEY | 92K | 189K | 585K | 296K | 233K | 649K | $6M |
| BANK OF AMERICA CORP /DE/ | 38K | 18K | 68K | 349K | 115K | 548K | $5M |
| GEODE CAPITAL MANAGEMENT, LLC | 334K | 360K | 416K | 455K | 461K | 525K | $5M |
Shares reported each quarter by the largest current holders. Green = added vs the prior filing, red = trimmed, grey = roughly held; a dash is a quarter with no 13F on file (a gap is skipped, not read as an exit). 13F is filed up to 45 days after quarter-end. Up to 40 quarters (10 years) available — use the selector above.
Line breaks are honest gaps — quarters with no 13F record, or a suppressed % — never interpolated. % of shares outstanding divides each quarter's as-filed 13F shares by the SEC cover-page share count dated to that same quarter (split-adjusted), not by today's count — so splits and buybacks don't move the line.
| Quarter | Holders | Shares held | % shares out | Value held |
|---|---|---|---|---|
| 2026-Q1 | 144 | 33,138,794 | 105.7% | $303M |
| 2025-Q4 | 161 | 35,070,814 | — | $562M |
| 2025-Q3 | 166 | 29,151,833 | 102.2% | $1.4B |
| 2025-Q2 | 114 | 24,772,291 | 88.0% | $726M |
| 2025-Q1 | 105 | 21,072,237 | 76.5% | $542M |
| 2024-Q4 | 95 | 19,863,850 | 72.2% | $573M |
| 2024-Q3 | 78 | 18,741,344 | 68.2% | $445M |
| 2024-Q2 | 65 | 18,918,982 | 74.1% | $327M |
| 2024-Q1 | 55 | 16,286,621 | — | $363M |
Short interest has its own tab now — open SI →
Source: SEC Form 13F — every institution manager (≥ $100M in 13F securities) reporting a long US-listed position, across ~10,250 filers over 10 years of history. 13F is long-only, US-listed, filed up to 45 days after quarter-end.
| Holder | % of class | Type | As of | |
|---|---|---|---|---|
| FMR LLC | 11.8% | QII | 2025-12-31 | SEC → |
| BlackRock, Inc. | 5.8% | QII | 2026-03-31 |
| Biggest buyers | Δ shares | Δ value | |
|---|---|---|---|
| VANGUARD CAPITAL MANAGEMENT LLC | +1,041,188 | $10M | OPEN |
| Squarepoint Ops LLC | +1,014,601 | $9M | ADD |
| Aberdeen Group plc | +666,729 | $6M | ADD |
| BANK OF AMERICA CORP /DE/ | +432,846 | $4M | ADD |
| MORGAN STANLEY | +415,466 | $4M | ADD |
| Biggest sellers | Δ shares | Δ value | |
|---|---|---|---|
| UBS Group AG | −1,414,749 | $13M | TRIM |
| VANGUARD GROUP INC | −1,321,202 | $12M | CLOSE |
| PRICE T ROWE ASSOCIATES INC /MD/ | −983,921 | $9M | TRIM |
| INTRINSIC EDGE CAPITAL MANAGEMENT LLC | −569,705 | $5M | CLOSE |
| Granahan Investment Management, LLC | −431,100 | $4M | CLOSE |
The 2025-Q4 register is as-filed in ITS OWN share basis; every count below was rebased into the 2026-Q1 basis (no split in between, factor 1). Dollar figures value the SHARE delta at one price — the 2026-03-31 close — rather than differencing two filings' own reported values, which would mix the quarter's price move into the trade figure.
13Fs disclose LONG US-listed positions only — shorts, most derivatives, bonds and non-US listings are invisible here, and a filer's quarter-over-quarter change is not necessarily a trade (index rebalances, in-kind transfers and entity successions all read as flow).
| SEC → |
| MILLENNIUM MANAGEMENT LLC | 5.1% | Passive | 2026-06-23 | SEC → |
| Squarepoint Ops LLC | 5.1% | Passive | 2026-04-01 | SEC → |
| Point72 Asset Management, L.P. | 4.8% | Passive | 2025-09-30 | SEC → |
Source: SEC Schedule 13D/13G structured XML. 13D = active intent; 13G filers report passively (Rule 13d-1(b) QII / 13d-1(c) Passive / 13d-1(d) Exempt) and are not activists. Percent and delta are the group aggregate the filing reports. Structured 13D/G coverage begins December 2024 (SEC XML mandate).
