KROS US Equity

Keros Therapeutics, Inc.

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Health Care · Pharmaceutical Preparations · CIK 1664710 · FYE 1231

$10.07

-0.14 (-1.37%)

USD · as of 2026-07-28 · marketstack

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Institutional ownership· 13F filers holding KROS · 2026-Q1 methodology → sources →

150 holders

% of shares outstanding held

49.1%

Shares held (13F)

18,595,183

Reporting holders

150

Top holders

largest 100 of 150 · Δ vs 2025-Q4

#	Holder	Shares	Value	% of shares out	Δ QoQ
1	BlackRock, Inc.	2,411,7932.4M	$27M	6.37%	ADD 5%
2	Nantahala Capital Management, LLC	1,360,4191.4M	$15M	3.59%	ADD 277%
3	VANGUARD CAPITAL MANAGEMENT LLC	831,421831,421	$9M	2.20%	NEW†
4	ORBIMED ADVISORS LLC	759,155759,155	$8M	2.01%	TRIM -6%
5	Qube Research & Technologies Ltd	724,926724,926	$8M	1.91%	TRIM -8%
6	JACOBS LEVY EQUITY MANAGEMENT, INC	706,071706,071	$8M	1.86%	TRIM -8%
7	FEDERATED HERMES, INC.	689,229689,229	$8M	1.82%	ADD 5%
8	VANGUARD PORTFOLIO MANAGEMENT LLC	682,043682,043	$8M	1.80%	NEW
9	GEODE CAPITAL MANAGEMENT, LLC	681,460681,460	$8M	1.80%	ADD 7%
10	Madison Avenue Partners, LP	626,583626,583	$7M	1.66%	TRIM -24%
11	ARROWSTREET CAPITAL, LIMITED PARTNERSHIP	590,249590,249	$7M	1.56%	TRIM -56%
12	STATE STREET CORP	561,730561,730	$6M	1.48%	TRIM -48%
13	D. E. Shaw & Co., Inc.	509,608509,608	$6M	1.35%	TRIM -33%
14	RENAISSANCE TECHNOLOGIES LLC	453,482453,482	$5M	1.20%	TRIM -23%
15	CITADEL ADVISORS LLC	406,666406,666	$4M	1.07%	TRIM -14%
16	AQR CAPITAL MANAGEMENT LLC	406,563406,563	$4M	1.07%	ADD 125%
17	TWO SIGMA INVESTMENTS, LP	363,835363,835	$4M	0.96%	ADD 14%
18	DIMENSIONAL FUND ADVISORS LP	321,360321,360	$4M	0.85%	ADD 454%
19	Nuveen, LLC	262,075262,075	$3M	0.69%	ADD 315%
20	Connor, Clark & Lunn Investment Management Ltd.	257,884257,884	$3M	0.68%	TRIM -20%
21	NORTHERN TRUST CORP	254,187254,187	$3M	0.67%	—
22	Man Group plc	238,618238,618	$3M	0.63%	ADD 25%
23	GOLDMAN SACHS GROUP INC	224,479224,479	$2M	0.59%	TRIM -62%
24	BRIDGEWAY CAPITAL MANAGEMENT, LLC	195,017195,017	$2M	0.52%	ADD 225%
25	CHARLES SCHWAB INVESTMENT MANAGEMENT INC	179,952179,952	$2M	0.48%	TRIM -4%
26	LOS ANGELES CAPITAL MANAGEMENT LLC	172,196172,196	$2M	0.45%	ADD 410%
27	Y-Intercept (Hong Kong) Ltd	164,483164,483	$2M	0.43%	ADD 40%
28	MACKENZIE FINANCIAL CORP	152,450152,450	$2M	0.40%	ADD 197%
29	WELLS FARGO & COMPANY/MN	151,040151,040	$2M	0.40%	ADD 361%
30	UBS Group AG	145,573145,573	$2M	0.38%	ADD 50%
31	Empowered Funds, LLC	145,517145,517	$2M	0.38%	NEW
32	BESSEMER GROUP INC	143,853143,853	$2M	0.38%	—
33	VANGUARD FIDUCIARY TRUST CO	127,302127,302	$1M	0.34%	NEW
34	MORGAN STANLEY	124,112124,112	$1M	0.33%	TRIM -46%
35	Bank of New York Mellon Corp	120,944120,944	$1M	0.32%	—
36	Point72 Asset Management, L.P.	109,015109,015	$1M	0.29%	ADD 181%
37	MILLENNIUM MANAGEMENT LLC	106,858106,858	$1M	0.28%	TRIM -51%
38	AMERIPRISE FINANCIAL INC	102,479102,479	$1M	0.27%	TRIM -17%
39	ADAR1 Capital Management, LLC	97,31897,318	$1M	0.26%	ADD 416%
40	XTX Topco Ltd	88,32288,322	$975,075	0.23%	ADD 747%
41	Invesco Ltd.	87,77287,772	$969,003	0.23%	—
42	Russell Investments Group, Ltd.	81,31181,311	$897,679	0.21%	—
43	HRT FINANCIAL LP	77,00277,002	$850,000	0.20%	TRIM -70%
44	ACADIAN ASSET MANAGEMENT LLC	65,43665,436	$721,000	0.17%	NEW
45	AMERICAN CENTURY COMPANIES INC	64,26064,260	$709,430	0.17%	ADD 75%
46	RBF Capital, LLC	62,80062,800	$693,312	0.17%	TRIM -62%
47	PRINCIPAL FINANCIAL GROUP INC	59,59459,594	$657,918	0.16%	NEW
48	ALLIANCEBERNSTEIN L.P.	30,86030,860	$628,310	0.08%	—
49	Rangeley Capital, LLC	54,40054,400	$600,576	0.14%	TRIM -44%
50	HSBC HOLDINGS PLC	53,98653,986	$598,705	0.14%	NEW
51	BANK OF AMERICA CORP /DE/	53,65453,654	$592,340	0.14%	TRIM -46%
52	DEUTSCHE BANK AG\	50,76550,765	$560,446	0.13%	TRIM -74%
53	STRS OHIO	50,60050,600	$558,624	0.13%	ADD 73%
54	Jump Financial, LLC	50,39350,393	$556,339	0.13%	TRIM -69%
55	PUBLIC EMPLOYEES RETIREMENT SYSTEM OF OHIO	50,32450,324	$555,577	0.13%	—
56	MASSACHUSETTS FINANCIAL SERVICES CO /MA/	48,99848,998	$540,938	0.13%	ADD 12%
57	MARTINGALE ASSET MANAGEMENT L P	47,59647,596	$525,460	0.13%	ADD 21%
58	PRELUDE CAPITAL MANAGEMENT, LLC	46,03946,039	$508,271	0.12%	ADD 238%
59	PRICE T ROWE ASSOCIATES INC /MD/	45,87645,876	$507,000	0.12%	ADD 54%
60	JANE STREET GROUP, LLC	44,45744,457	$490,805	0.12%	TRIM -73%
61	CANADA PENSION PLAN INVESTMENT BOARD	44,10044,100	$486,864	0.12%	—
62	RHUMBLINE ADVISERS	40,17640,176	$443,530	0.11%	TRIM -9%
63	GLOBEFLEX CAPITAL L P	37,87337,873	$418,118	0.10%	NEW
64	JPMORGAN CHASE & CO	37,68037,680	$402,423	0.10%	ADD 102%
65	WINTON GROUP Ltd	36,43336,433	$402,220	0.10%	ADD 69%
66	PANAGORA ASSET MANAGEMENT INC	34,65634,656	$382,602	0.09%	TRIM -72%
67	AVIVA PLC	34,30734,307	$378,749	0.09%	NEW
68	PDT Partners, LLC	32,74432,744	$361,494	0.09%	TRIM -71%
69	FMR LLC	32,67432,674	$360,721	0.09%	TRIM -8%
70	Magnetar Financial LLC	30,08530,085	$332,138	0.08%	NEW
71	TUDOR INVESTMENT CORP ET AL	28,00728,007	$309,197	0.07%	—
72	Quantinno Capital Management LP	23,75923,759	$262,299	0.06%	ADD 12%
73	BARCLAYS PLC	23,15723,157	$255,652	0.06%	TRIM -69%
74	SIMPLEX TRADING, LLC	22,28822,288	$246,059	0.06%	NEW
75	FOX RUN MANAGEMENT, L.L.C.	22,79022,790	$243,397	0.06%	TRIM -11%
76	FRANKLIN RESOURCES INC	21,75621,756	$240,186	0.06%	TRIM -21%
77	LSV ASSET MANAGEMENT	20,90020,900	$231,000	0.06%	TRIM -10%
78	Creative Planning	17,80717,807	$196,589	0.05%	TRIM -23%
79	MetLife Investment Management, LLC	16,68216,682	$184,169	0.04%	—
80	Engineers Gate Manager LP	16,59516,595	$183,209	0.04%	TRIM -73%
81	STATE OF WISCONSIN INVESTMENT BOARD	15,93215,932	$175,889	0.04%	TRIM -20%
82	Vanguard Global Advisers, LLC	15,75515,755	$173,935	0.04%	NEW
83	EJF Capital LP	15,00015,000	$165,600	0.04%	—
84	Brevan Howard Capital Management LP	14,65114,651	$161,747	0.04%	NEW
85	MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.	13,89113,891	$153,357	0.04%	TRIM -9%
86	Mariner, LLC	13,82613,826	$152,616	0.04%	TRIM -9%
87	Virtus Investment Advisers, LLC	12,67612,676	$139,943	0.03%	ADD 2%
88	MERCER GLOBAL ADVISORS INC /ADV	11,74211,742	$129,632	0.03%	NEW
89	LAZARD ASSET MANAGEMENT LLC	11,70511,705	$129,223	0.03%	NEW
90	SUSQUEHANNA INTERNATIONAL GROUP, LLP	11,24211,242	$124,112	0.03%	TRIM -16%
91	OXFORD ASSET MANAGEMENT LLP	11,15311,153	$123,129	0.03%	TRIM -48%
92	AMERICAN FINANCIAL GROUP INC	10,10510,105	$111,559	0.03%	—
93	SG Americas Securities, LLC	9,1359,135	$101,000	0.02%	TRIM -91%
94	NEW YORK STATE COMMON RETIREMENT FUND	9,1169,116	$100,641	0.02%	TRIM -14%
95	CWM, LLC	8,6188,618	$95,143	0.02%	ADD 125%
96	Police & Firemen's Retirement System of New Jersey	7,7007,700	$85,008	0.02%	—
97	BNP PARIBAS FINANCIAL MARKETS	6,9046,904	$76,220	0.02%	ADD 60%
98	China Universal Asset Management Co., Ltd.	5,4065,406	$59,682	0.01%	ADD 9%
99	ROYAL BANK OF CANADA	4,8674,867	$54,000	0.01%	TRIM -14%
100	Sterling Capital Management LLC	4,3614,361	$48,145	0.01%	TRIM -38%

† a filer this large appearing as NEW usually reflects a renamed or succeeded filing entity (the old name stops filing the same quarter) — not a brand-new position.

Rows rank by the value each filer reported — 13F filers price their own positions, so a holder with fewer shares can rank just above one with more.

Holder trends· top 15 holders · shares held over 6 quarters

green added · red trimmed

Holder

2024-Q4

2025-Q1

2025-Q2

2025-Q3

2025-Q4

2026-Q1

Latest value

BlackRock, Inc.

2.7M

2.8M

2.7M

2.3M

2.4M

$27M

Nantahala Capital Management, LLC

915K

1.5M

1.6M

1.2M

360K

1.4M

$15M

VANGUARD CAPITAL MANAGEMENT LLC

—

831K

$9M

ORBIMED ADVISORS LLC

1.0M

804K

759K

$8M

Qube Research & Technologies Ltd

—

273K

39K

303K

784K

725K

$8M

JACOBS LEVY EQUITY MANAGEMENT, INC

—

524K

607K

768K

706K

$8M

FEDERATED HERMES, INC.

125K

1.2M

654K

514K

655K

689K

$8M

VANGUARD PORTFOLIO MANAGEMENT LLC

—

682K

$8M

GEODE CAPITAL MANAGEMENT, LLC

765K

806K

829K

837K

635K

681K

$8M

Madison Avenue Partners, LP

—

2.7M

824K

627K

$7M

ARROWSTREET CAPITAL, LIMITED PARTNERSHIP

—

65K

259K

1.3M

590K

$7M

STATE STREET CORP

1.2M

1.3M

1.4M

1.3M

1.1M

562K

$6M

D. E. Shaw & Co., Inc.

12K

1.3M

755K

510K

$6M

RENAISSANCE TECHNOLOGIES LLC

—

147K

155K

592K

453K

$5M

CITADEL ADVISORS LLC

791K

192K

23K

176K

473K

407K

$4M

Shares reported each quarter by the largest current holders. Green = added vs the prior filing, red = trimmed, grey = roughly held; a dash is a quarter with no 13F on file (a gap is skipped, not read as an exit). 13F is filed up to 45 days after quarter-end. Up to 40 quarters (10 years) available — use the selector above.

Institutional footprint by quarter

Line breaks are honest gaps — quarters with no 13F record, or a suppressed % — never interpolated. % of shares outstanding uses today's share count for every quarter (the only count stored).

Quarter	Holders	Shares held	% shares out	Value held
2026-Q1	150	18,595,183	49.1%	$206M
2025-Q4	165	20,156,272	53.2%	$411M
2025-Q3	155	35,452,711	93.6%	$561M
2025-Q2	139	35,940,128	94.9%	$480M
2025-Q1	156	36,741,531	97.0%	$429M
2024-Q4	161	36,764,494	97.1%	$582M
2024-Q3	164	34,986,611	92.4%	$2.0B
2024-Q2	152	31,907,135	84.3%	$1.5B
2024-Q1	147	32,034,262	84.6%	$2.1B
2023-Q4	121

Short interest has its own tab now — open SI →

Source: SEC Form 13F — every institution manager (≥ $100M in 13F securities) reporting a long US-listed position, across ~10,250 filers over 10 years of history. 13F is long-only, US-listed, filed up to 45 days after quarter-end.

Activists & large holders· Schedule 13D/G beneficial owners · KROS

Activists — 13D, active intent

Pontifax Management 4 G.P. (2015) Ltd.0%EXITEDSCHEDULE 13D/A · Amd 2 · event 2025-10-15SEC →

0 shares · 6 reporting persons

“Item 4 is hereby amended and supplemented with the addition of the following: On October 15, 2025, certain of the Reporting Persons (collectively, the "Pontifax Parties") and the Issuer entered into a Stock Purchase Agreement (the "Repurchase Agreement"). Pursuant to the Repurchase Agreement, the Pontifax Parties agreed to sell a total of 4,787,331 shares of Common Stock, representing all the shares of Common Stock owned by the Reporting Persons, at a per share price of $17.75, for an aggregate ”

ADAR1 Capital Management, LLC0%EXITEDSCHEDULE 13D/A · Amd 5 · event 2025-10-15SEC →

0 shares · 3 reporting persons

“Item 4 of the Schedule 13D is hereby amended and supplemented by the addition of the following: On October 15, 2025, ADAR 1 Partners, LP, ADAR1 SPV I, LP and Spearhead Insurance Solutions IDF, LLC (the "ADAR1 Funds") and ADAR1 Capital Management (together with the ADAR1 Funds, the "ADAR1 Parties") entered into a stock purchase agreement (the "Stock Purchase Agreement") with the Issuer pursuant to which the ADAR1 Funds agreed to sell a total of 5,389,264 shares of Common Stock to the Issuer at a ”

Large holders — 13G, passive (top 5 of 11 by % of class)

Institutional ownership· 13F filers holding KROS · 2026-Q1 methodology → sources →

150 holders

% of shares outstanding held

49.1%

Shares held (13F)

18,595,183

Reporting holders

150

Top holders

largest 100 of 150 · Δ vs 2025-Q4

#	Holder	Shares	Value	% of shares out	Δ QoQ
1	BlackRock, Inc.	2,411,7932.4M	$27M	6.37%	ADD 5%
2	Nantahala Capital Management, LLC	1,360,4191.4M	$15M	3.59%	ADD 277%
3	VANGUARD CAPITAL MANAGEMENT LLC	831,421831,421	$9M	2.20%	NEW†
4	ORBIMED ADVISORS LLC	759,155759,155	$8M	2.01%	TRIM -6%
5	Qube Research & Technologies Ltd	724,926724,926	$8M	1.91%	TRIM -8%
6	JACOBS LEVY EQUITY MANAGEMENT, INC	706,071706,071	$8M	1.86%	TRIM -8%
7	FEDERATED HERMES, INC.	689,229689,229	$8M	1.82%	ADD 5%
8	VANGUARD PORTFOLIO MANAGEMENT LLC	682,043682,043	$8M	1.80%	NEW
9	GEODE CAPITAL MANAGEMENT, LLC	681,460681,460	$8M	1.80%	ADD 7%
10	Madison Avenue Partners, LP	626,583626,583	$7M	1.66%	TRIM -24%
11	ARROWSTREET CAPITAL, LIMITED PARTNERSHIP	590,249590,249	$7M	1.56%	TRIM -56%
12	STATE STREET CORP	561,730561,730	$6M	1.48%	TRIM -48%
13	D. E. Shaw & Co., Inc.	509,608509,608	$6M	1.35%	TRIM -33%
14	RENAISSANCE TECHNOLOGIES LLC	453,482453,482	$5M	1.20%	TRIM -23%
15	CITADEL ADVISORS LLC	406,666406,666	$4M	1.07%	TRIM -14%
16	AQR CAPITAL MANAGEMENT LLC	406,563406,563	$4M	1.07%	ADD 125%
17	TWO SIGMA INVESTMENTS, LP	363,835363,835	$4M	0.96%	ADD 14%
18	DIMENSIONAL FUND ADVISORS LP	321,360321,360	$4M	0.85%	ADD 454%
19	Nuveen, LLC	262,075262,075	$3M	0.69%	ADD 315%
20	Connor, Clark & Lunn Investment Management Ltd.	257,884257,884	$3M	0.68%	TRIM -20%
21	NORTHERN TRUST CORP	254,187254,187	$3M	0.67%	—
22	Man Group plc	238,618238,618	$3M	0.63%	ADD 25%
23	GOLDMAN SACHS GROUP INC	224,479224,479	$2M	0.59%	TRIM -62%
24	BRIDGEWAY CAPITAL MANAGEMENT, LLC	195,017195,017	$2M	0.52%	ADD 225%
25	CHARLES SCHWAB INVESTMENT MANAGEMENT INC	179,952179,952	$2M	0.48%	TRIM -4%
26	LOS ANGELES CAPITAL MANAGEMENT LLC	172,196172,196	$2M	0.45%	ADD 410%
27	Y-Intercept (Hong Kong) Ltd	164,483164,483	$2M	0.43%	ADD 40%
28	MACKENZIE FINANCIAL CORP	152,450152,450	$2M	0.40%	ADD 197%
29	WELLS FARGO & COMPANY/MN	151,040151,040	$2M	0.40%	ADD 361%
30	UBS Group AG	145,573145,573	$2M	0.38%	ADD 50%
31	Empowered Funds, LLC	145,517145,517	$2M	0.38%	NEW
32	BESSEMER GROUP INC	143,853143,853	$2M	0.38%	—
33	VANGUARD FIDUCIARY TRUST CO	127,302127,302	$1M	0.34%	NEW
34	MORGAN STANLEY	124,112124,112	$1M	0.33%	TRIM -46%
35	Bank of New York Mellon Corp	120,944120,944	$1M	0.32%	—
36	Point72 Asset Management, L.P.	109,015109,015	$1M	0.29%	ADD 181%
37	MILLENNIUM MANAGEMENT LLC	106,858106,858	$1M	0.28%	TRIM -51%
38	AMERIPRISE FINANCIAL INC	102,479102,479	$1M	0.27%	TRIM -17%
39	ADAR1 Capital Management, LLC	97,31897,318	$1M	0.26%	ADD 416%
40	XTX Topco Ltd	88,32288,322	$975,075	0.23%	ADD 747%
41	Invesco Ltd.	87,77287,772	$969,003	0.23%	—
42	Russell Investments Group, Ltd.	81,31181,311	$897,679	0.21%	—
43	HRT FINANCIAL LP	77,00277,002	$850,000	0.20%	TRIM -70%
44	ACADIAN ASSET MANAGEMENT LLC	65,43665,436	$721,000	0.17%	NEW
45	AMERICAN CENTURY COMPANIES INC	64,26064,260	$709,430	0.17%	ADD 75%
46	RBF Capital, LLC	62,80062,800	$693,312	0.17%	TRIM -62%
47	PRINCIPAL FINANCIAL GROUP INC	59,59459,594	$657,918	0.16%	NEW
48	ALLIANCEBERNSTEIN L.P.	30,86030,860	$628,310	0.08%	—
49	Rangeley Capital, LLC	54,40054,400	$600,576	0.14%	TRIM -44%
50	HSBC HOLDINGS PLC	53,98653,986	$598,705	0.14%	NEW
51	BANK OF AMERICA CORP /DE/	53,65453,654	$592,340	0.14%	TRIM -46%
52	DEUTSCHE BANK AG\	50,76550,765	$560,446	0.13%	TRIM -74%
53	STRS OHIO	50,60050,600	$558,624	0.13%	ADD 73%
54	Jump Financial, LLC	50,39350,393	$556,339	0.13%	TRIM -69%
55	PUBLIC EMPLOYEES RETIREMENT SYSTEM OF OHIO	50,32450,324	$555,577	0.13%	—
56	MASSACHUSETTS FINANCIAL SERVICES CO /MA/	48,99848,998	$540,938	0.13%	ADD 12%
57	MARTINGALE ASSET MANAGEMENT L P	47,59647,596	$525,460	0.13%	ADD 21%
58	PRELUDE CAPITAL MANAGEMENT, LLC	46,03946,039	$508,271	0.12%	ADD 238%
59	PRICE T ROWE ASSOCIATES INC /MD/	45,87645,876	$507,000	0.12%	ADD 54%
60	JANE STREET GROUP, LLC	44,45744,457	$490,805	0.12%	TRIM -73%
61	CANADA PENSION PLAN INVESTMENT BOARD	44,10044,100	$486,864	0.12%	—
62	RHUMBLINE ADVISERS	40,17640,176	$443,530	0.11%	TRIM -9%
63	GLOBEFLEX CAPITAL L P	37,87337,873	$418,118	0.10%	NEW
64	JPMORGAN CHASE & CO	37,68037,680	$402,423	0.10%	ADD 102%
65	WINTON GROUP Ltd	36,43336,433	$402,220	0.10%	ADD 69%
66	PANAGORA ASSET MANAGEMENT INC	34,65634,656	$382,602	0.09%	TRIM -72%
67	AVIVA PLC	34,30734,307	$378,749	0.09%	NEW
68	PDT Partners, LLC	32,74432,744	$361,494	0.09%	TRIM -71%
69	FMR LLC	32,67432,674	$360,721	0.09%	TRIM -8%
70	Magnetar Financial LLC	30,08530,085	$332,138	0.08%	NEW
71	TUDOR INVESTMENT CORP ET AL	28,00728,007	$309,197	0.07%	—
72	Quantinno Capital Management LP	23,75923,759	$262,299	0.06%	ADD 12%
73	BARCLAYS PLC	23,15723,157	$255,652	0.06%	TRIM -69%
74	SIMPLEX TRADING, LLC	22,28822,288	$246,059	0.06%	NEW
75	FOX RUN MANAGEMENT, L.L.C.	22,79022,790	$243,397	0.06%	TRIM -11%
76	FRANKLIN RESOURCES INC	21,75621,756	$240,186	0.06%	TRIM -21%
77	LSV ASSET MANAGEMENT	20,90020,900	$231,000	0.06%	TRIM -10%
78	Creative Planning	17,80717,807	$196,589	0.05%	TRIM -23%
79	MetLife Investment Management, LLC	16,68216,682	$184,169	0.04%	—
80	Engineers Gate Manager LP	16,59516,595	$183,209	0.04%	TRIM -73%
81	STATE OF WISCONSIN INVESTMENT BOARD	15,93215,932	$175,889	0.04%	TRIM -20%
82	Vanguard Global Advisers, LLC	15,75515,755	$173,935	0.04%	NEW
83	EJF Capital LP	15,00015,000	$165,600	0.04%	—
84	Brevan Howard Capital Management LP	14,65114,651	$161,747	0.04%	NEW
85	MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.	13,89113,891	$153,357	0.04%	TRIM -9%
86	Mariner, LLC	13,82613,826	$152,616	0.04%	TRIM -9%
87	Virtus Investment Advisers, LLC	12,67612,676	$139,943	0.03%	ADD 2%
88	MERCER GLOBAL ADVISORS INC /ADV	11,74211,742	$129,632	0.03%	NEW
89	LAZARD ASSET MANAGEMENT LLC	11,70511,705	$129,223	0.03%	NEW
90	SUSQUEHANNA INTERNATIONAL GROUP, LLP	11,24211,242	$124,112	0.03%	TRIM -16%
91	OXFORD ASSET MANAGEMENT LLP	11,15311,153	$123,129	0.03%	TRIM -48%
92	AMERICAN FINANCIAL GROUP INC	10,10510,105	$111,559	0.03%	—
93	SG Americas Securities, LLC	9,1359,135	$101,000	0.02%	TRIM -91%
94	NEW YORK STATE COMMON RETIREMENT FUND	9,1169,116	$100,641	0.02%	TRIM -14%
95	CWM, LLC	8,6188,618	$95,143	0.02%	ADD 125%
96	Police & Firemen's Retirement System of New Jersey	7,7007,700	$85,008	0.02%	—
97	BNP PARIBAS FINANCIAL MARKETS	6,9046,904	$76,220	0.02%	ADD 60%
98	China Universal Asset Management Co., Ltd.	5,4065,406	$59,682	0.01%	ADD 9%
99	ROYAL BANK OF CANADA	4,8674,867	$54,000	0.01%	TRIM -14%
100	Sterling Capital Management LLC	4,3614,361	$48,145	0.01%	TRIM -38%

† a filer this large appearing as NEW usually reflects a renamed or succeeded filing entity (the old name stops filing the same quarter) — not a brand-new position.

Rows rank by the value each filer reported — 13F filers price their own positions, so a holder with fewer shares can rank just above one with more.

Holder trends· top 15 holders · shares held over 6 quarters

green added · red trimmed

Holder

2024-Q4

2025-Q1

2025-Q2

2025-Q3

2025-Q4

2026-Q1

Latest value

BlackRock, Inc.

2.7M

2.8M

2.7M

2.3M

2.4M

$27M

Nantahala Capital Management, LLC

915K

1.5M

1.6M

1.2M

360K

1.4M

$15M

VANGUARD CAPITAL MANAGEMENT LLC

—

831K

$9M

ORBIMED ADVISORS LLC

1.0M

804K

759K

$8M

Qube Research & Technologies Ltd

—

273K

39K

303K

784K

725K

$8M

JACOBS LEVY EQUITY MANAGEMENT, INC

—

524K

607K

768K

706K

$8M

FEDERATED HERMES, INC.

125K

1.2M

654K

514K

655K

689K

$8M

VANGUARD PORTFOLIO MANAGEMENT LLC

—

682K

$8M

GEODE CAPITAL MANAGEMENT, LLC

765K

806K

829K

837K

635K

681K

$8M

Madison Avenue Partners, LP

—

2.7M

824K

627K

$7M

ARROWSTREET CAPITAL, LIMITED PARTNERSHIP

—

65K

259K

1.3M

590K

$7M

STATE STREET CORP

1.2M

1.3M

1.4M

1.3M

1.1M

562K

$6M

D. E. Shaw & Co., Inc.

12K

1.3M

755K

510K

$6M

RENAISSANCE TECHNOLOGIES LLC

—

147K

155K

592K

453K

$5M

CITADEL ADVISORS LLC

791K

192K

23K

176K

473K

407K

$4M

Institutional footprint by quarter

Line breaks are honest gaps — quarters with no 13F record, or a suppressed % — never interpolated. % of shares outstanding uses today's share count for every quarter (the only count stored).

Quarter	Holders	Shares held	% shares out	Value held
2026-Q1	150	18,595,183	49.1%	$206M
2025-Q4	165	20,156,272	53.2%	$411M
2025-Q3	155	35,452,711	93.6%	$561M
2025-Q2	139	35,940,128	94.9%	$480M
2025-Q1	156	36,741,531	97.0%	$429M
2024-Q4	161	36,764,494	97.1%	$582M
2024-Q3	164	34,986,611	92.4%	$2.0B
2024-Q2	152	31,907,135	84.3%	$1.5B
2024-Q1	147	32,034,262	84.6%	$2.1B
2023-Q4	121

Short interest has its own tab now — open SI →

Activists & large holders· Schedule 13D/G beneficial owners · KROS

Activists — 13D, active intent

Pontifax Management 4 G.P. (2015) Ltd.0%EXITEDSCHEDULE 13D/A · Amd 2 · event 2025-10-15SEC →

0 shares · 6 reporting persons

ADAR1 Capital Management, LLC0%EXITEDSCHEDULE 13D/A · Amd 5 · event 2025-10-15SEC →

0 shares · 3 reporting persons

Large holders — 13G, passive (top 5 of 11 by % of class)

Holder	% of class	Type	As of
BlackRock, Inc.	10.1%	QII	2026-03-31	SEC →
Nantahala Capital Management, LLC	6.9%	QII	2026-03-31	SEC →
BlackRock Portfolio Management LLC	5.7%	QII	2026-06-30	SEC →
D. E. SHAW & CO, L.P.	3.9%	Passive	2025-12-31	SEC →
Madison Avenue Partners, LP	2.7%	Passive	2025-12-31	SEC →

KROS US Equity☆

Institutional ownership· 13F filers holding KROS · 2026-Q1 methodology → sources →

Top holders

Holder trends· top 15 holders · shares held over 6 quarters

Institutional footprint by quarter

Activists & large holders· Schedule 13D/G beneficial owners · KROS

Institutional ownership· 13F filers holding KROS · 2026-Q1 methodology → sources →

Top holders

Holder trends· top 15 holders · shares held over 6 quarters

Institutional footprint by quarter

Activists & large holders· Schedule 13D/G beneficial owners · KROS

KROS US Equity