CDNA US Equity

CareDx, Inc.

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Health Care · Services-Medical Laboratories · CIK 1217234 · FYE 1231

$35.41

-0.34 (-0.95%)

USD · as of 2026-07-29 · marketstack

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Institutional ownership· 13F filers holding CDNA · 2026-Q1 methodology → sources →

190 holders

% of shares outstanding held

101.7%

Shares held (13F)

52,532,510

Reporting holders

190

Institutional 13F holdings sum to 101.7% of shares outstanding. Readings above 100% reflect securities-lending double counting: shares on loan to short sellers are reported long by both the lending institution and the one that bought them. FINRA short interest of 6,892,213 shares (settlement 2026-04-30) accounts for the excess.

Top holders

largest 100 of 190 · Δ vs 2025-Q4

#	Holder	Shares	Value	% of shares out	Δ QoQ
1	BlackRock, Inc.	4,522,0964.5M	$79M	8.75%	TRIM -3%
2	BAMCO INC /NY/	3,336,4093.3M	$58M	6.46%	ADD 6%
3	STATE STREET CORP	2,760,3852.8M	$48M	5.34%	—
4	ARK Investment Management LLC	2,227,5562.2M	$39M	4.31%	ADD 4%
5	DEERFIELD MANAGEMENT COMPANY, L.P.	2,191,0002.2M	$38M	4.24%	—
6	BRAIDWELL LP	2,173,1462.2M	$38M	4.21%	ADD 37%
7	VANGUARD CAPITAL MANAGEMENT LLC	2,160,4572.2M	$38M	4.18%	NEW†
8	Boston Partners	1,883,5341.9M	$33M	3.65%	ADD 109%
9	GAGNON SECURITIES LLC	1,761,8741.8M	$31M	3.41%	—
10	Chevy Chase Trust Holdings, LLC	1,504,0751.5M	$26M	2.91%	ADD 39%
11	Sumitomo Mitsui Trust Group, Inc.	1,475,1261.5M	$26M	2.86%	—
12	Amova Asset Management Americas, Inc.	1,475,1261.5M	$26M	2.86%	—
13	DIMENSIONAL FUND ADVISORS LP	1,461,9141.5M	$25M	2.83%	ADD 12%
14	GEODE CAPITAL MANAGEMENT, LLC	1,254,1141.3M	$22M	2.43%	—
15	RENAISSANCE TECHNOLOGIES LLC	1,184,3981.2M	$21M	2.29%	ADD 7%
16	Ophir Asset Management Pty Ltd	988,051988,051	$17M	1.91%	ADD 7%
17	Blue Water Life Science Advisors, LP	935,000935,000	$16M	1.81%	ADD 83%
18	D. E. Shaw & Co., Inc.	874,908874,908	$15M	1.69%	TRIM -16%
19	JACOBS LEVY EQUITY MANAGEMENT, INC	864,468864,468	$15M	1.67%	TRIM -9%
20	VANGUARD PORTFOLIO MANAGEMENT LLC	851,433851,433	$15M	1.65%	NEW
21	GOLDMAN SACHS GROUP INC	733,332733,332	$13M	1.42%	TRIM -10%
22	First Eagle Investment Management, LLC	575,365575,365	$10M	1.11%	ADD 5%
23	Russell Investments Group, Ltd.	558,336558,336	$10M	1.08%	ADD 33%
24	ROYCE & ASSOCIATES LP	553,197553,197	$10M	1.07%	TRIM -5%
25	Gagnon Advisors, LLC	532,892532,892	$9M	1.03%	—
26	Palo Alto Investors LP	531,870531,870	$9M	1.03%	ADD 11%
27	CHARLES SCHWAB INVESTMENT MANAGEMENT INC	510,405510,405	$9M	0.99%	TRIM -3%
28	PARAGON ASSOCIATES & PARAGON ASSOCIATES II JOINT VENTURE	500,000500,000	$9M	0.97%	ADD 33%
29	ArrowMark Colorado Holdings LLC	486,668486,668	$8M	0.94%	TRIM -36%
30	Invesco Ltd.	484,938484,938	$8M	0.94%	ADD 320%
31	BANK OF AMERICA CORP /DE/	470,665470,665	$8M	0.91%	ADD 45%
32	BNP PARIBAS FINANCIAL MARKETS	458,301458,301	$8M	0.89%	TRIM -6%
33	NORTHERN TRUST CORP	454,508454,508	$8M	0.88%	—
34	MORGAN STANLEY	425,349425,349	$7M	0.82%	TRIM -21%
35	Aberdeen Group plc	417,744417,744	$7M	0.81%	ADD 21%
36	Nuveen, LLC	390,294390,294	$7M	0.76%	ADD 17%
37	Zweig-DiMenna Associates LLC	390,255390,255	$7M	0.76%	TRIM -43%
38	Connor, Clark & Lunn Investment Management Ltd.	370,183370,183	$6M	0.72%	TRIM -44%
39	VANGUARD FIDUCIARY TRUST CO	352,466352,466	$6M	0.68%	NEW
40	AQR CAPITAL MANAGEMENT LLC	349,225349,225	$6M	0.68%	ADD 15%
41	Portolan Capital Management, LLC	340,352340,352	$6M	0.66%	TRIM -55%
42	Meridiem Capital Partners LP	316,950316,950	$6M	0.61%	NEW
43	JANUS HENDERSON GROUP PLC	316,519316,519	$5M	0.61%	ADD 10%
44	SILVERCREST ASSET MANAGEMENT GROUP LLC	310,539310,539	$5M	0.60%	TRIM -16%
45	Parkman Healthcare Partners LLC	291,378291,378	$5M	0.56%	ADD 6%
46	JANE STREET GROUP, LLC	250,351250,351	$4M	0.48%	TRIM -42%
47	TWO SIGMA INVESTMENTS, LP	231,600231,600	$4M	0.45%	ADD 159%
48	FMR LLC	231,305231,305	$4M	0.45%	ADD 10%
49	Bullseye Asset Management LLC	220,485220,485	$4M	0.43%	TRIM -6%
50	Legal & General Group Plc	209,763209,763	$4M	0.41%	TRIM -9%
51	Schonfeld Strategic Advisors LLC	197,925197,925	$3M	0.38%	ADD 151%
52	FRED ALGER MANAGEMENT, LLC	173,013173,013	$3M	0.33%	—
53	Bank of New York Mellon Corp	170,347170,347	$3M	0.33%	—
54	Man Group plc	166,474166,474	$3M	0.32%	TRIM -29%
55	RAYMOND JAMES FINANCIAL INC	156,483156,483	$3M	0.30%	TRIM -3%
56	WELLINGTON MANAGEMENT GROUP LLP	145,680145,680	$3M	0.28%	TRIM -14%
57	STATE BOARD OF ADMINISTRATION OF FLORIDA RETIREMENT SYSTEM	139,439139,439	$2M	0.27%	—
58	GLOBEFLEX CAPITAL L P	119,252119,252	$2M	0.23%	ADD 49%
59	LAZARD ASSET MANAGEMENT LLC	116,751116,751	$2M	0.23%	—
60	UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC	115,785115,785	$2M	0.22%	TRIM -8%
61	Engineers Gate Manager LP	115,207115,207	$2M	0.22%	TRIM -4%
62	AMERICAN CENTURY COMPANIES INC	114,350114,350	$2M	0.22%	ADD 31%
63	IQ EQ FUND MANAGEMENT (IRELAND) Ltd	101,747101,747	$2M	0.20%	ADD 22%
64	Swiss National Bank	99,70099,700	$2M	0.19%	—
65	STATE OF WISCONSIN INVESTMENT BOARD	98,31798,317	$2M	0.19%	TRIM -21%
66	UBS Group AG	94,15994,159	$2M	0.18%	TRIM -22%
67	Gotham Asset Management, LLC	89,60489,604	$2M	0.17%	ADD 8%
68	CITADEL ADVISORS LLC	84,17384,173	$1M	0.16%	TRIM -87%
69	Rafferty Asset Management, LLC	78,73478,734	$1M	0.15%	TRIM -14%
70	FORMULA GROWTH LTD	78,47878,478	$1M	0.15%	TRIM -22%
71	ARS Investment Partners, LLC	77,23577,235	$1M	0.15%	TRIM -18%
72	FRANKLIN RESOURCES INC	69,44469,444	$1M	0.13%	TRIM -33%
73	PRINCIPAL FINANCIAL GROUP INC	68,92568,925	$1M	0.13%	TRIM -12%
74	RHUMBLINE ADVISERS	65,50065,500	$1M	0.13%	TRIM -5%
75	Uniplan Investment Counsel, Inc.	62,77362,773	$1M	0.12%	ADD 122%
76	MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.	62,67362,673	$1M	0.12%	—
77	PRUDENTIAL FINANCIAL INC	60,24560,245	$1M	0.12%	ADD 9%
78	AMERIPRISE FINANCIAL INC	51,90751,907	$901,106	0.10%	—
79	SUSQUEHANNA INTERNATIONAL GROUP, LLP	50,64650,646	$879,215	0.10%	ADD 7%
80	ALLIANCEBERNSTEIN L.P.	46,46046,460	$875,306	0.09%	—
81	DEUTSCHE BANK AG\	49,87349,873	$865,795	0.10%	ADD 10%
82	PRICE T ROWE ASSOCIATES INC /MD/	47,00147,001	$816,000	0.09%	ADD 20%
83	CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM	45,90345,903	$796,876	0.09%	—
84	ALGERT GLOBAL LLC	44,51044,510	$773,000	0.09%	TRIM -37%
85	EXCHANGE TRADED CONCEPTS, LLC	44,29644,296	$768,979	0.09%	—
86	Y-Intercept (Hong Kong) Ltd	41,29441,294	$716,864	0.08%	ADD 59%
87	VIRTUS ADVISERS, LLC	40,56440,564	$704,191	0.08%	—
88	CANADA PENSION PLAN INVESTMENT BOARD	40,10040,100	$696,136	0.08%	ADD 4%
89	JPMORGAN CHASE & CO	40,32340,323	$662,507	0.08%	ADD 2%
90	HSBC HOLDINGS PLC	38,38738,387	$644,134	0.07%	ADD 15%
91	Palumbo Wealth Management LLC	36,41036,410	$632,078	0.07%	TRIM -20%
92	PROFUND ADVISORS LLC	35,21735,217	$611,367	0.07%	ADD 4%
93	VANGUARD ASSET MANAGEMENT, Ltd	33,23233,232	$576,908	0.06%	NEW
94	Creative Planning	30,49730,497	$529,428	0.06%	ADD 3%
95	SG Americas Securities, LLC	29,94929,949	$520,000	0.06%	ADD 8%
96	MetLife Investment Management, LLC	29,02129,021	$503,805	0.06%	—
97	CITIGROUP INC	28,19128,191	$489,395	0.05%	ADD 3%
98	XTX Topco Ltd	27,72027,720	$481,219	0.05%	NEW
99	State of Wyoming	23,53723,537	$408,602	0.05%	ADD 185%
100	Allspring Global Investments Holdings, LLC	21,24221,242	$379,595	0.04%	ADD 11%

† a filer this large appearing as NEW usually reflects a renamed or succeeded filing entity (the old name stops filing the same quarter) — not a brand-new position.

Rows rank by the value each filer reported — 13F filers price their own positions, so a holder with fewer shares can rank just above one with more.

Holder trends· top 15 holders · shares held over 6 quarters

green added · red trimmed

Holder

2024-Q4

2025-Q1

2025-Q2

2025-Q3

2025-Q4

2026-Q1

Latest value

BlackRock, Inc.

5.1M

4.7M

4.8M

4.7M

4.5M

$79M

BAMCO INC /NY/

2.2M

2.4M

3.2M

3.3M

$58M

STATE STREET CORP

2.7M

2.5M

2.2M

2.6M

2.7M

2.8M

$48M

ARK Investment Management LLC

2.4M

2.3M

2.1M

2.2M

2.1M

2.2M

$39M

DEERFIELD MANAGEMENT COMPANY, L.P.

—

2.2M

$38M

BRAIDWELL LP

—

2.2M

1.6M

2.2M

$38M

VANGUARD CAPITAL MANAGEMENT LLC

—

2.2M

$38M

Boston Partners

21K

31K

902K

1.9M

$33M

GAGNON SECURITIES LLC

1.4M

1.5M

1.7M

1.8M

$31M

Chevy Chase Trust Holdings, LLC

—

535K

513K

744K

1.1M

1.5M

$26M

Sumitomo Mitsui Trust Group, Inc.

1.5M

1.4M

1.2M

1.7M

1.4M

1.5M

$26M

Amova Asset Management Americas, Inc.

1.5M

1.4M

1.2M

1.7M

1.4M

1.5M

$26M

DIMENSIONAL FUND ADVISORS LP

743K

692K

963K

1.2M

1.3M

1.5M

$25M

GEODE CAPITAL MANAGEMENT, LLC

1.2M

1.3M

1.2M

1.3M

$22M

RENAISSANCE TECHNOLOGIES LLC

1.4M

1.2M

1.1M

1.2M

$21M

Shares reported each quarter by the largest current holders. Green = added vs the prior filing, red = trimmed, grey = roughly held; a dash is a quarter with no 13F on file (a gap is skipped, not read as an exit). 13F is filed up to 45 days after quarter-end. Up to 40 quarters (10 years) available — use the selector above.

Institutional footprint by quarter

Line breaks are honest gaps — quarters with no 13F record, or a suppressed % — never interpolated. % of shares outstanding uses today's share count for every quarter (the only count stored).

Quarter	Holders	Shares held	% shares out	Value held
2026-Q1	190	52,532,510	101.7%	$912M
2025-Q4	200	52,179,231	101.0%	$983M
2025-Q3	187	53,635,628	103.8%

Short interest has its own tab now — open SI →

Source: SEC Form 13F — every institution manager (≥ $100M in 13F securities) reporting a long US-listed position, across ~10,250 filers over 10 years of history. 13F is long-only, US-listed, filed up to 45 days after quarter-end.

Activists & large holders· Schedule 13D/G beneficial owners · CDNA

Large holders — 13G, passive (top 5 of 6 by % of class)

Holder	% of class	Type	As of
BlackRock, Inc.	9%	QII	2026-06-30	SEC →
BAMCO INC /NY/	5.93%	QII	2025-09-30	SEC →
GAGNON SECURITIES LLC	5.9%	Passive	2025-12-31	SEC →
STATE STREET CORP	5.3%	QII	2025-12-31	SEC →
ARK Investment Management LLC	4.49%	QII	2024-12-31

Institutional ownership· 13F filers holding CDNA · 2026-Q1 methodology → sources →

190 holders

% of shares outstanding held

101.7%

Shares held (13F)

52,532,510

Reporting holders

190

Top holders

largest 100 of 190 · Δ vs 2025-Q4

#	Holder	Shares	Value	% of shares out	Δ QoQ
1	BlackRock, Inc.	4,522,0964.5M	$79M	8.75%	TRIM -3%
2	BAMCO INC /NY/	3,336,4093.3M	$58M	6.46%	ADD 6%
3	STATE STREET CORP	2,760,3852.8M	$48M	5.34%	—
4	ARK Investment Management LLC	2,227,5562.2M	$39M	4.31%	ADD 4%
5	DEERFIELD MANAGEMENT COMPANY, L.P.	2,191,0002.2M	$38M	4.24%	—
6	BRAIDWELL LP	2,173,1462.2M	$38M	4.21%	ADD 37%
7	VANGUARD CAPITAL MANAGEMENT LLC	2,160,4572.2M	$38M	4.18%	NEW†
8	Boston Partners	1,883,5341.9M	$33M	3.65%	ADD 109%
9	GAGNON SECURITIES LLC	1,761,8741.8M	$31M	3.41%	—
10	Chevy Chase Trust Holdings, LLC	1,504,0751.5M	$26M	2.91%	ADD 39%
11	Sumitomo Mitsui Trust Group, Inc.	1,475,1261.5M	$26M	2.86%	—
12	Amova Asset Management Americas, Inc.	1,475,1261.5M	$26M	2.86%	—
13	DIMENSIONAL FUND ADVISORS LP	1,461,9141.5M	$25M	2.83%	ADD 12%
14	GEODE CAPITAL MANAGEMENT, LLC	1,254,1141.3M	$22M	2.43%	—
15	RENAISSANCE TECHNOLOGIES LLC	1,184,3981.2M	$21M	2.29%	ADD 7%
16	Ophir Asset Management Pty Ltd	988,051988,051	$17M	1.91%	ADD 7%
17	Blue Water Life Science Advisors, LP	935,000935,000	$16M	1.81%	ADD 83%
18	D. E. Shaw & Co., Inc.	874,908874,908	$15M	1.69%	TRIM -16%
19	JACOBS LEVY EQUITY MANAGEMENT, INC	864,468864,468	$15M	1.67%	TRIM -9%
20	VANGUARD PORTFOLIO MANAGEMENT LLC	851,433851,433	$15M	1.65%	NEW
21	GOLDMAN SACHS GROUP INC	733,332733,332	$13M	1.42%	TRIM -10%
22	First Eagle Investment Management, LLC	575,365575,365	$10M	1.11%	ADD 5%
23	Russell Investments Group, Ltd.	558,336558,336	$10M	1.08%	ADD 33%
24	ROYCE & ASSOCIATES LP	553,197553,197	$10M	1.07%	TRIM -5%
25	Gagnon Advisors, LLC	532,892532,892	$9M	1.03%	—
26	Palo Alto Investors LP	531,870531,870	$9M	1.03%	ADD 11%
27	CHARLES SCHWAB INVESTMENT MANAGEMENT INC	510,405510,405	$9M	0.99%	TRIM -3%
28	PARAGON ASSOCIATES & PARAGON ASSOCIATES II JOINT VENTURE	500,000500,000	$9M	0.97%	ADD 33%
29	ArrowMark Colorado Holdings LLC	486,668486,668	$8M	0.94%	TRIM -36%
30	Invesco Ltd.	484,938484,938	$8M	0.94%	ADD 320%
31	BANK OF AMERICA CORP /DE/	470,665470,665	$8M	0.91%	ADD 45%
32	BNP PARIBAS FINANCIAL MARKETS	458,301458,301	$8M	0.89%	TRIM -6%
33	NORTHERN TRUST CORP	454,508454,508	$8M	0.88%	—
34	MORGAN STANLEY	425,349425,349	$7M	0.82%	TRIM -21%
35	Aberdeen Group plc	417,744417,744	$7M	0.81%	ADD 21%
36	Nuveen, LLC	390,294390,294	$7M	0.76%	ADD 17%
37	Zweig-DiMenna Associates LLC	390,255390,255	$7M	0.76%	TRIM -43%
38	Connor, Clark & Lunn Investment Management Ltd.	370,183370,183	$6M	0.72%	TRIM -44%
39	VANGUARD FIDUCIARY TRUST CO	352,466352,466	$6M	0.68%	NEW
40	AQR CAPITAL MANAGEMENT LLC	349,225349,225	$6M	0.68%	ADD 15%
41	Portolan Capital Management, LLC	340,352340,352	$6M	0.66%	TRIM -55%
42	Meridiem Capital Partners LP	316,950316,950	$6M	0.61%	NEW
43	JANUS HENDERSON GROUP PLC	316,519316,519	$5M	0.61%	ADD 10%
44	SILVERCREST ASSET MANAGEMENT GROUP LLC	310,539310,539	$5M	0.60%	TRIM -16%
45	Parkman Healthcare Partners LLC	291,378291,378	$5M	0.56%	ADD 6%
46	JANE STREET GROUP, LLC	250,351250,351	$4M	0.48%	TRIM -42%
47	TWO SIGMA INVESTMENTS, LP	231,600231,600	$4M	0.45%	ADD 159%
48	FMR LLC	231,305231,305	$4M	0.45%	ADD 10%
49	Bullseye Asset Management LLC	220,485220,485	$4M	0.43%	TRIM -6%
50	Legal & General Group Plc	209,763209,763	$4M	0.41%	TRIM -9%
51	Schonfeld Strategic Advisors LLC	197,925197,925	$3M	0.38%	ADD 151%
52	FRED ALGER MANAGEMENT, LLC	173,013173,013	$3M	0.33%	—
53	Bank of New York Mellon Corp	170,347170,347	$3M	0.33%	—
54	Man Group plc	166,474166,474	$3M	0.32%	TRIM -29%
55	RAYMOND JAMES FINANCIAL INC	156,483156,483	$3M	0.30%	TRIM -3%
56	WELLINGTON MANAGEMENT GROUP LLP	145,680145,680	$3M	0.28%	TRIM -14%
57	STATE BOARD OF ADMINISTRATION OF FLORIDA RETIREMENT SYSTEM	139,439139,439	$2M	0.27%	—
58	GLOBEFLEX CAPITAL L P	119,252119,252	$2M	0.23%	ADD 49%
59	LAZARD ASSET MANAGEMENT LLC	116,751116,751	$2M	0.23%	—
60	UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC	115,785115,785	$2M	0.22%	TRIM -8%
61	Engineers Gate Manager LP	115,207115,207	$2M	0.22%	TRIM -4%
62	AMERICAN CENTURY COMPANIES INC	114,350114,350	$2M	0.22%	ADD 31%
63	IQ EQ FUND MANAGEMENT (IRELAND) Ltd	101,747101,747	$2M	0.20%	ADD 22%
64	Swiss National Bank	99,70099,700	$2M	0.19%	—
65	STATE OF WISCONSIN INVESTMENT BOARD	98,31798,317	$2M	0.19%	TRIM -21%
66	UBS Group AG	94,15994,159	$2M	0.18%	TRIM -22%
67	Gotham Asset Management, LLC	89,60489,604	$2M	0.17%	ADD 8%
68	CITADEL ADVISORS LLC	84,17384,173	$1M	0.16%	TRIM -87%
69	Rafferty Asset Management, LLC	78,73478,734	$1M	0.15%	TRIM -14%
70	FORMULA GROWTH LTD	78,47878,478	$1M	0.15%	TRIM -22%
71	ARS Investment Partners, LLC	77,23577,235	$1M	0.15%	TRIM -18%
72	FRANKLIN RESOURCES INC	69,44469,444	$1M	0.13%	TRIM -33%
73	PRINCIPAL FINANCIAL GROUP INC	68,92568,925	$1M	0.13%	TRIM -12%
74	RHUMBLINE ADVISERS	65,50065,500	$1M	0.13%	TRIM -5%
75	Uniplan Investment Counsel, Inc.	62,77362,773	$1M	0.12%	ADD 122%
76	MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.	62,67362,673	$1M	0.12%	—
77	PRUDENTIAL FINANCIAL INC	60,24560,245	$1M	0.12%	ADD 9%
78	AMERIPRISE FINANCIAL INC	51,90751,907	$901,106	0.10%	—
79	SUSQUEHANNA INTERNATIONAL GROUP, LLP	50,64650,646	$879,215	0.10%	ADD 7%
80	ALLIANCEBERNSTEIN L.P.	46,46046,460	$875,306	0.09%	—
81	DEUTSCHE BANK AG\	49,87349,873	$865,795	0.10%	ADD 10%
82	PRICE T ROWE ASSOCIATES INC /MD/	47,00147,001	$816,000	0.09%	ADD 20%
83	CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM	45,90345,903	$796,876	0.09%	—
84	ALGERT GLOBAL LLC	44,51044,510	$773,000	0.09%	TRIM -37%
85	EXCHANGE TRADED CONCEPTS, LLC	44,29644,296	$768,979	0.09%	—
86	Y-Intercept (Hong Kong) Ltd	41,29441,294	$716,864	0.08%	ADD 59%
87	VIRTUS ADVISERS, LLC	40,56440,564	$704,191	0.08%	—
88	CANADA PENSION PLAN INVESTMENT BOARD	40,10040,100	$696,136	0.08%	ADD 4%
89	JPMORGAN CHASE & CO	40,32340,323	$662,507	0.08%	ADD 2%
90	HSBC HOLDINGS PLC	38,38738,387	$644,134	0.07%	ADD 15%
91	Palumbo Wealth Management LLC	36,41036,410	$632,078	0.07%	TRIM -20%
92	PROFUND ADVISORS LLC	35,21735,217	$611,367	0.07%	ADD 4%
93	VANGUARD ASSET MANAGEMENT, Ltd	33,23233,232	$576,908	0.06%	NEW
94	Creative Planning	30,49730,497	$529,428	0.06%	ADD 3%
95	SG Americas Securities, LLC	29,94929,949	$520,000	0.06%	ADD 8%
96	MetLife Investment Management, LLC	29,02129,021	$503,805	0.06%	—
97	CITIGROUP INC	28,19128,191	$489,395	0.05%	ADD 3%
98	XTX Topco Ltd	27,72027,720	$481,219	0.05%	NEW
99	State of Wyoming	23,53723,537	$408,602	0.05%	ADD 185%
100	Allspring Global Investments Holdings, LLC	21,24221,242	$379,595	0.04%	ADD 11%

† a filer this large appearing as NEW usually reflects a renamed or succeeded filing entity (the old name stops filing the same quarter) — not a brand-new position.

Rows rank by the value each filer reported — 13F filers price their own positions, so a holder with fewer shares can rank just above one with more.

Holder trends· top 15 holders · shares held over 6 quarters

green added · red trimmed

Holder

2024-Q4

2025-Q1

2025-Q2

2025-Q3

2025-Q4

2026-Q1

Latest value

BlackRock, Inc.

5.1M

4.7M

4.8M

4.7M

4.5M

$79M

BAMCO INC /NY/

2.2M

2.4M

3.2M

3.3M

$58M

STATE STREET CORP

2.7M

2.5M

2.2M

2.6M

2.7M

2.8M

$48M

ARK Investment Management LLC

2.4M

2.3M

2.1M

2.2M

2.1M

2.2M

$39M

DEERFIELD MANAGEMENT COMPANY, L.P.

—

2.2M

$38M

BRAIDWELL LP

—

2.2M

1.6M

2.2M

$38M

VANGUARD CAPITAL MANAGEMENT LLC

—

2.2M

$38M

Boston Partners

21K

31K

902K

1.9M

$33M

GAGNON SECURITIES LLC

1.4M

1.5M

1.7M

1.8M

$31M

Chevy Chase Trust Holdings, LLC

—

535K

513K

744K

1.1M

1.5M

$26M

Sumitomo Mitsui Trust Group, Inc.

1.5M

1.4M

1.2M

1.7M

1.4M

1.5M

$26M

Amova Asset Management Americas, Inc.

1.5M

1.4M

1.2M

1.7M

1.4M

1.5M

$26M

DIMENSIONAL FUND ADVISORS LP

743K

692K

963K

1.2M

1.3M

1.5M

$25M

GEODE CAPITAL MANAGEMENT, LLC

1.2M

1.3M

1.2M

1.3M

$22M

RENAISSANCE TECHNOLOGIES LLC

1.4M

1.2M

1.1M

1.2M

$21M

Institutional footprint by quarter

Line breaks are honest gaps — quarters with no 13F record, or a suppressed % — never interpolated. % of shares outstanding uses today's share count for every quarter (the only count stored).

Quarter	Holders	Shares held	% shares out	Value held
2026-Q1	190	52,532,510	101.7%	$912M
2025-Q4	200	52,179,231	101.0%	$983M
2025-Q3	187	53,635,628	103.8%

Short interest has its own tab now — open SI →

Activists & large holders· Schedule 13D/G beneficial owners · CDNA

Large holders — 13G, passive (top 5 of 6 by % of class)

Holder	% of class	Type	As of
BlackRock, Inc.	9%	QII	2026-06-30	SEC →
BAMCO INC /NY/	5.93%	QII	2025-09-30	SEC →
GAGNON SECURITIES LLC	5.9%	Passive	2025-12-31	SEC →
STATE STREET CORP	5.3%	QII	2025-12-31	SEC →
ARK Investment Management LLC	4.49%	QII	2024-12-31

CDNA US Equity☆

Institutional ownership· 13F filers holding CDNA · 2026-Q1 methodology → sources →

Top holders

Holder trends· top 15 holders · shares held over 6 quarters

Institutional footprint by quarter

Activists & large holders· Schedule 13D/G beneficial owners · CDNA

Institutional ownership· 13F filers holding CDNA · 2026-Q1 methodology → sources →

Top holders

Holder trends· top 15 holders · shares held over 6 quarters

Institutional footprint by quarter

Activists & large holders· Schedule 13D/G beneficial owners · CDNA

CDNA US Equity