| # | Holder | Shares | Value | % of shares out | Δ QoQ |
|---|---|---|---|---|---|
| 1 | ADAMS STREET PARTNERS LLC | 1,711,4301.7M | $135M | 8.06% | — |
| 2 | BAILLIE GIFFORD & CO | 1,259,9931.3M | $99M | 5.93% | — |
| 3 | Civilization Ventures Management, LLC | 1,099,4311.1M | $87M | 5.18% | — |
| 4 | BAMCO INC /NY/ | 741,485741,485 | $59M | 3.49% | ADD 336% |
| 5 | PRICE T ROWE ASSOCIATES INC /MD/ | 737,518737,518 | $58M | 3.47% | ADD 52% |
| 6 | NVP ASSOCIATES, LLC | 513,330513,330 | $41M | 2.42% | — |
| 7 | NEUBERGER BERMAN GROUP LLC | 709,214709,214 | $38M | 3.34% | — |
| 8 | Invesco Ltd. | 406,512406,512 | $32M | 1.91% | ADD 114% |
| 9 | WASATCH ADVISORS LP | 369,638369,638 | $29M | 1.74% | NEW |
| 10 | JPMORGAN CHASE & CO | 283,140283,140 | $20M | 1.33% | TRIM -28% |
| 11 | UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC | 240,587240,587 | $19M | 1.13% | ADD 6% |
| 12 | EMERALD ADVISERS, LLC | 239,562239,562 | $19M | 1.13% | ADD 27% |
| 13 | Napean Trading & Investment Co (Singapore) PTE Ltd | 233,333233,333 | $18M | 1.10% | — |
| 14 | ALLIANCEBERNSTEIN L.P. | 223,251223,251 | $18M | 1.05% | ADD 7% |
| 15 | AMERIPRISE FINANCIAL INC | 220,035220,035 | $17M | 1.04% | ADD 80% |
| 16 | VANGUARD CAPITAL MANAGEMENT LLC | 182,463182,463 | $14M | 0.86% | NEW |
| 17 | Capital International Investors | 178,000178,000 | $14M | 0.84% | — |
| 18 | MASSACHUSETTS FINANCIAL SERVICES CO /MA/ | 159,573159,573 | $13M | 0.75% | TRIM -2% |
| 19 | VANGUARD PORTFOLIO MANAGEMENT LLC | 151,213151,213 | $12M | 0.71% | NEW |
| 20 | EMERALD MUTUAL FUND ADVISERS TRUST | 132,206132,206 | $10M | 0.62% | ADD 12% |
| 21 | Penn Capital Management Company, LLC | 126,335126,335 | $10M | 0.60% | NEW |
| 22 | UBS Group AG | 121,604121,604 | $10M | 0.57% | ADD 285% |
| 23 | BlackRock, Inc. | 95,16095,160 | $8M | 0.45% | ADD 33% |
| 24 | PERCEPTIVE ADVISORS LLC | 91,81291,812 | $7M | 0.43% | ADD 26% |
| 25 | NEXT CENTURY GROWTH INVESTORS LLC | 90,56490,564 | $7M | 0.43% | ADD 147% |
| 26 | GEODE CAPITAL MANAGEMENT, LLC | 89,11489,114 | $7M | 0.42% | ADD 9% |
| 27 | NEW YORK STATE COMMON RETIREMENT FUND | 88,68288,682 | $7M | 0.42% | ADD 1633% |
| 28 | PFM Health Sciences, LP | 88,06988,069 | $7M | 0.41% | — |
| 29 | TWO SIGMA INVESTMENTS, LP | 82,77682,776 | $7M | 0.39% | NEW |
| 30 | Ensign Peak Advisors, Inc | 51,50151,501 | $4M | 0.24% | NEW |
| 31 | RENAISSANCE TECHNOLOGIES LLC | 51,25451,254 | $4M | 0.24% | NEW |
| 32 | Deep Track Capital, LP | 50,00050,000 | $4M | 0.24% | — |
| 33 | LORD, ABBETT & CO. LLC | 44,28944,289 | $3M | 0.21% | TRIM -4% |
| 34 | TD Asset Management Inc | 43,10843,108 | $3M | 0.20% | ADD 61% |
| 35 | MANUFACTURERS LIFE INSURANCE COMPANY, THE | 30,50630,506 | $2M | 0.14% | ADD 152% |
| 36 | CHARLES SCHWAB INVESTMENT MANAGEMENT INC | 28,05428,054 | $2M | 0.13% | ADD 5% |
| 37 | VANGUARD FIDUCIARY TRUST CO | 27,79127,791 | $2M | 0.13% | NEW |
| 38 | SEVEN GRAND MANAGERS, LLC | 25,00025,000 | $2M | 0.12% | — |
| 39 | Squarepoint Ops LLC | 22,94622,946 | $2M | 0.11% | ADD 191% |
| 40 | Universal- Beteiligungs- und Servicegesellschaft mbH | 20,59520,595 | $2M | 0.10% | ADD 25% |
| 41 | Blue Water Life Science Advisors, LP | 20,00020,000 | $2M | 0.09% | NEW |
| 42 | STATE STREET CORP | 19,70019,700 | $2M | 0.09% | TRIM -9% |
| 43 | PUBLIC EMPLOYEES RETIREMENT SYSTEM OF OHIO | 19,49919,499 | $2M | 0.09% | NEW |
| 44 | Russell Investments Group, Ltd. | 18,65918,659 | $1M | 0.09% | NEW |
| 45 | F/m Investments LLC | 16,90816,908 | $1M | 0.08% | NEW |
| 46 | SOROS FUND MANAGEMENT LLC | 14,50014,500 | $1M | 0.07% | — |
| 47 | DEUTSCHE BANK AG\ | 13,64013,640 | $1M | 0.06% | NEW |
| 48 | SEI INVESTMENTS CO | 13,39913,399 | $1M | 0.06% | NEW |
| 49 | XTX Topco Ltd | 13,17913,179 | $1M | 0.06% | TRIM -36% |
| 50 | BANK OF AMERICA CORP /DE/ | 12,70712,707 | $1M | 0.06% | ADD 8603% |
| 51 | MORGAN STANLEY | 12,58612,586 | $993,539 | 0.06% | ADD 250% |
| 52 | SUSQUEHANNA INTERNATIONAL GROUP, LLP | 12,48612,486 | $985,645 | 0.06% | NEW |
| 53 | Bellevue Group AG | 10,55410,554 | $833,133 | 0.05% | — |
| 54 | NOMURA HOLDINGS INC | 8,6008,600 | $678,884 | 0.04% | NEW |
| 55 | Brevan Howard Capital Management LP | 7,7167,716 | $609,101 | 0.04% | ADD 69% |
| 56 | Alyeska Investment Group, L.P. | 7,0007,000 | $552,580 | 0.03% | — |
| 57 | Walleye Capital LLC | 5,9565,956 | $470,167 | 0.03% | — |
| 58 | BRANDYWINE MANAGERS, LLC | 5,7505,750 | $453,905 | 0.03% | NEW |
| 59 | Green Alpha Advisors, LLC | 5,1685,168 | $407,962 | 0.02% | TRIM -3% |
| 60 | ROYAL BANK OF CANADA | 4,7704,770 | $377,000 | 0.02% | TRIM -14% |
| 61 | Pathstone Holdings, LLC | 4,2804,280 | $337,864 | 0.02% | — |
| 62 | GOLDMAN SACHS GROUP INC | 4,1584,158 | $328,233 | 0.02% | TRIM -80% |
| 63 | MYDA Advisors LLC | 4,0004,000 | $315,760 | 0.02% | TRIM -90% |
| 64 | D. E. Shaw & Co., Inc. | 3,8623,862 | $304,866 | 0.02% | NEW |
| 65 | CONNABLE OFFICE INC | 3,8503,850 | $303,919 | 0.02% | NEW |
| 66 | Cornerstone Investment Partners, LLC | 3,4023,402 | $268,554 | 0.02% | NEW |
| 67 | BARCLAYS PLC | 2,7582,758 | $217,717 | 0.01% | NEW |
| 68 | Renaissance Capital LLC | 1,9271,927 | $144,621 | 0.01% | TRIM -21% |
| 69 | Vanguard Global Advisers, LLC | 1,7551,755 | $138,540 | 0.01% | NEW |
| 70 | Smartleaf Asset Management LLC | 1,2971,297 | $90,933 | 0.01% | ADD 38% |
| 71 | WELLS FARGO & COMPANY/MN | 1,1081,108 | $87,466 | 0.01% | ADD 19% |
| 72 | SBI Securities Co., Ltd. | 680680 | $53,679 | 0.00% | ADD 858% |
| 73 | RiverPark Advisors, LLC | 627627 | $49,495 | 0.00% | ADD 35% |
| 74 | OSAIC HOLDINGS, INC. | 420420 | $33,154 | 0.00% | ADD 35% |
| 75 | Steward Partners Investment Advisory, LLC | 300300 | $23,682 | 0.00% | ADD 200% |
| 76 | HARBOUR INVESTMENTS, INC. | 180180 | $14,209 | 0.00% | NEW |
| 77 | FIFTH THIRD BANCORP | 134134 | $10,615 | 0.00% | NEW |
| 78 | Rossby Financial, LCC | 100100 | $8,296 | 0.00% | TRIM -67% |
| 79 | MCF Advisors LLC | 104104 | $8,210 | 0.00% | ADD 167% |
| 80 | US BANCORP \DE\ | 6464 | $5,052 | 0.00% | NEW |
| 81 | Colonial Trust Advisors | 5050 | $3,947 | 0.00% | — |
| 82 | Quantbot Technologies LP | 4040 | $3,158 | 0.00% | NEW |
| 83 | Physician Wealth Advisors, Inc. | 1717 | $1,342 | 0.00% | — |
| 84 | HUNTINGTON NATIONAL BANK | 1515 | $1,184 | 0.00% | NEW |
| 85 | Frazier Financial Advisors, LLC | 1414 | $1,106 | 0.00% | — |
| 86 | JONES FINANCIAL COMPANIES LLLP | 1212 | $835 | 0.00% | — |
| 87 | BNP PARIBAS FINANCIAL MARKETS | 88 | $632 | 0.00% | NEW |
| 88 | Marquette Asset Management, LLC | 77 | $553 | 0.00% | NEW |
| 89 | CITIGROUP INC | 11 | $79 | 0.00% | — |
Rows rank by the value each filer reported — 13F filers price their own positions, so a holder with fewer shares can rank just above one with more.
| Holder | 2024-Q4 | 2025-Q1 | 2025-Q2 | 2025-Q3 | 2025-Q4 | 2026-Q1 | Latest value |
|---|---|---|---|---|---|---|---|
| ADAMS STREET PARTNERS LLC | — | — | — | — | 1.7M | 1.7M | $135M |
| BAILLIE GIFFORD & CO | — | — | — | — | 1.3M | 1.3M | $99M |
| Civilization Ventures Management, LLC | — | — | — | — | 1.1M | 1.1M | $87M |
| BAMCO INC /NY/ | — | — | — | — | 170K | 741K | $59M |
| PRICE T ROWE ASSOCIATES INC /MD/ | — | — | — | — | 485K | 738K | $58M |
| NVP ASSOCIATES, LLC | — | — | — | — | 513K | 513K | $41M |
| NEUBERGER BERMAN GROUP LLC | — | — | — | — | 701K | 709K | $38M |
| Invesco Ltd. | — | — | — | — | 190K | 407K | $32M |
| WASATCH ADVISORS LP | — | — | — | — | — | 370K | $29M |
| JPMORGAN CHASE & CO | — | — | — | — | 392K | 283K | $20M |
| UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC | — | — | — | — | 228K | 241K | $19M |
| EMERALD ADVISERS, LLC | — | — | — | — | 189K | 240K | $19M |
| Napean Trading & Investment Co (Singapore) PTE Ltd | — | — | — | — | 233K | 233K | $18M |
| ALLIANCEBERNSTEIN L.P. | — | — | — | — | 209K | 223K | $18M |
| AMERIPRISE FINANCIAL INC | — | — | — | — | 122K | 220K | $17M |
Shares reported each quarter by the largest current holders. Green = added vs the prior filing, red = trimmed, grey = roughly held; a dash is a quarter with no 13F on file (a gap is skipped, not read as an exit). 13F is filed up to 45 days after quarter-end. Up to 40 quarters (10 years) available — use the selector above.
Line breaks are honest gaps — quarters with no 13F record, or a suppressed % — never interpolated. % of shares outstanding uses today's share count for every quarter (the only count stored).
| Quarter | Holders | Shares held | % shares out | Value held |
|---|---|---|---|---|
| 2026-Q1 | 89 | 11,339,021 | 53.4% | $875M |
| 2025-Q4 | 89 | 9,651,838 | 45.5% | $790M |
Short interest has its own tab now — open SI →
Source: SEC Form 13F — every institution manager (≥ $100M in 13F securities) reporting a long US-listed position, across ~10,250 filers over 10 years of history. 13F is long-only, US-listed, filed up to 45 days after quarter-end.
