VCYT US Equity

VERACYTE, INC.

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Health Care · Services-Medical Laboratories · CIK 1384101 · FYE 1231

$55.65

+0.04 (+0.07%)

USD · as of 2026-07-28 · marketstack

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Institutional ownership· 13F filers holding VCYT · 2026-Q1 methodology → sources →

334 holders

% of shares outstanding held

110.6%

Shares held (13F)

88,283,972

Reporting holders

334

Institutional 13F holdings sum to 110.6% of shares outstanding. Readings above 100% reflect securities-lending double counting: shares on loan to short sellers are reported long by both the lending institution and the one that bought them. FINRA short interest of 6,870,870 shares (settlement 2026-04-30) accounts for the excess.

Top holders

largest 100 of 334 · Δ vs 2025-Q4

#	Holder	Shares	Value	% of shares out	Δ QoQ
1	BlackRock, Inc.	13,172,61813.2M	$424M	16.51%	—
2	FMR LLC	7,763,8497.8M	$250M	9.73%	ADD 35%
3	Artisan Partners Limited Partnership	5,496,3705.5M	$177M	6.89%	TRIM -12%
4	STATE STREET CORP	5,043,5905.0M	$162M	6.32%	ADD 16%
5	VANGUARD PORTFOLIO MANAGEMENT LLC	4,444,7304.4M	$143M	5.57%	NEW†
6	VANGUARD CAPITAL MANAGEMENT LLC	3,532,2323.5M	$114M	4.43%	NEW†
7	ARK Investment Management LLC	3,053,8183.1M	$98M	3.83%	TRIM -21%
8	DIMENSIONAL FUND ADVISORS LP	2,881,0902.9M	$93M	3.61%	ADD 3%
9	WELLINGTON MANAGEMENT GROUP LLP	2,558,2852.6M	$82M	3.21%	ADD 26%
10	FIRST TRUST ADVISORS LP	2,117,0462.1M	$68M	2.65%	ADD 65%
11	GEODE CAPITAL MANAGEMENT, LLC	1,957,0962.0M	$63M	2.45%	ADD 3%
12	BANK OF AMERICA CORP /DE/	1,448,8971.4M	$47M	1.82%	ADD 15%
13	GW&K Investment Management, LLC	1,437,3221.4M	$46M	1.80%	ADD 26%
14	FULLER & THALER ASSET MANAGEMENT, INC.	1,414,3621.4M	$46M	1.77%	TRIM -49%
15	ArrowMark Colorado Holdings LLC	1,374,1031.4M	$44M	1.72%	TRIM -12%
16	GOLDMAN SACHS GROUP INC	1,276,2601.3M	$41M	1.60%	ADD 7%
17	Sumitomo Mitsui Trust Group, Inc.	1,055,6561.1M	$34M	1.32%	TRIM -16%
18	Amova Asset Management Americas, Inc.	1,045,0471.0M	$34M	1.31%	TRIM -17%
19	WILLIAM BLAIR INVESTMENT MANAGEMENT, LLC	989,687989,687	$32M	1.24%	ADD 25%
20	NORTHERN TRUST CORP	929,500929,500	$30M	1.16%	TRIM -6%
21	CHARLES SCHWAB INVESTMENT MANAGEMENT INC	923,573923,573	$30M	1.16%	—
22	Legal & General Group Plc	897,171897,171	$29M	1.12%	ADD 30%
23	OBERWEIS ASSET MANAGEMENT INC/	839,834839,834	$27M	1.05%	ADD 16%
24	Alyeska Investment Group, L.P.	808,570808,570	$26M	1.01%	ADD 34%
25	MORGAN STANLEY	797,802797,802	$26M	1.00%	TRIM -33%
26	Fisher Asset Management, LLC	736,270736,270	$24M	0.92%	TRIM -7%
27	DEERFIELD MANAGEMENT COMPANY, L.P.	726,000726,000	$23M	0.91%	—
28	BAMCO INC /NY/	655,522655,522	$21M	0.82%	—
29	SEGALL BRYANT & HAMILL, LLC	591,488591,488	$19M	0.74%	—
30	FRED ALGER MANAGEMENT, LLC	579,460579,460	$19M	0.73%	TRIM -9%
31	ROYAL BANK OF CANADA	573,096573,096	$18M	0.72%	ADD 66%
32	VANGUARD FIDUCIARY TRUST CO	548,167548,167	$18M	0.69%	NEW
33	Point72 Asset Management, L.P.	483,904483,904	$16M	0.61%	ADD 14%
34	PRINCIPAL FINANCIAL GROUP INC	476,686476,686	$15M	0.60%	ADD 2%
35	JPMORGAN CHASE & CO	496,219496,219	$15M	0.62%	TRIM -5%
36	Invesco Ltd.	436,587436,587	$14M	0.55%	ADD 12%
37	NEW YORK STATE COMMON RETIREMENT FUND	421,633421,633	$14M	0.53%	TRIM -28%
38	SEI INVESTMENTS CO	420,093420,093	$14M	0.53%	ADD 9%
39	Bank of New York Mellon Corp	413,416413,416	$13M	0.52%	TRIM -3%
40	BNP PARIBAS FINANCIAL MARKETS	362,836362,836	$12M	0.45%	ADD 3%
41	JUPITER ASSET MANAGEMENT LTD	355,604355,604	$11M	0.45%	—
42	RAYMOND JAMES FINANCIAL INC	370,261370,261	$11M	0.46%	TRIM -4%
43	AEGON ASSET MANAGEMENT UK Plc	326,627326,627	$11M	0.41%	TRIM -33%
44	Soleus Capital Management, L.P.	319,912319,912	$10M	0.40%	ADD 3099%
45	Balyasny Asset Management L.P.	298,955298,955	$10M	0.37%	NEW
46	Granite Investment Partners, LLC	298,589298,589	$10M	0.37%	TRIM -15%
47	Public Sector Pension Investment Board	282,835282,835	$9M	0.35%	ADD 3%
48	Champlain Investment Partners, LLC	281,923281,923	$9M	0.35%	TRIM -59%
49	Casdin Capital, LLC	276,308276,308	$9M	0.35%	—
50	Nuveen, LLC	238,439238,439	$8M	0.30%	TRIM -59%
51	Summit Creek Advisors LLC	225,894225,894	$7M	0.28%	TRIM -5%
52	RENAISSANCE TECHNOLOGIES LLC	223,500223,500	$7M	0.28%	NEW
53	NORDEA INVESTMENT MANAGEMENT AB	227,581227,581	$7M	0.29%	TRIM -4%
54	RHUMBLINE ADVISERS	213,234213,234	$7M	0.27%	—
55	FRANKLIN RESOURCES INC	211,683211,683	$7M	0.27%	—
56	UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC	205,462205,462	$7M	0.26%	ADD 4%
57	BLAIR WILLIAM & CO/IL	194,887194,887	$6M	0.24%	—
58	PRUDENTIAL FINANCIAL INC	190,910190,910	$6M	0.24%	ADD 28%
59	JANUS HENDERSON GROUP PLC	187,368187,368	$6M	0.23%	TRIM -31%
60	SILVERARC CAPITAL MANAGEMENT, LLC	179,700179,700	$6M	0.23%	TRIM -50%
61	Man Group plc	177,876177,876	$6M	0.22%	ADD 1430%
62	UBS Group AG	172,496172,496	$6M	0.22%	—
63	Palisades Investment Partners, LLC	170,603170,603	$5M	0.21%	ADD 5%
64	Allianz Asset Management GmbH	158,871158,871	$5M	0.20%	ADD 17%
65	Swiss National Bank	156,400156,400	$5M	0.20%	—
66	CITADEL ADVISORS LLC	152,941152,941	$5M	0.19%	TRIM -4%
67	NEUBERGER BERMAN GROUP LLC	150,805150,805	$5M	0.19%	ADD 2876%
68	EXCHANGE TRADED CONCEPTS, LLC	145,925145,925	$5M	0.18%	ADD 32%
69	ARROWSTREET CAPITAL, LIMITED PARTNERSHIP	141,811141,811	$5M	0.18%	NEW
70	AMERIPRISE FINANCIAL INC	135,073135,073	$4M	0.17%	TRIM -5%
71	ALLIANCEBERNSTEIN L.P.	102,643102,643	$4M	0.13%	—
72	Vestal Point Capital, LP	130,687130,687	$4M	0.16%	NEW
73	Trexquant Investment LP	130,159130,159	$4M	0.16%	ADD 1556%
74	Corient Private Wealth LLC	127,722127,722	$4M	0.16%	TRIM -51%
75	EFG International AG	125,345125,345	$4M	0.16%	TRIM -6%
76	ARDSLEY ADVISORY PARTNERS LP	125,000125,000	$4M	0.16%	—
77	IQ EQ FUND MANAGEMENT (IRELAND) Ltd	124,331124,331	$4M	0.16%	TRIM -20%
78	BROWN CAPITAL MANAGEMENT LLC	107,819107,819	$3M	0.14%	TRIM -6%
79	PINNACLE ASSOCIATES LTD	107,769107,769	$3M	0.14%	ADD 105%
80	RICE HALL JAMES & ASSOCIATES, LLC	106,665106,665	$3M	0.13%	—
81	Campbell & CO Investment Adviser LLC	106,443106,443	$3M	0.13%	ADD 581%
82	Rafferty Asset Management, LLC	104,212104,212	$3M	0.13%	ADD 21%
83	MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.	102,445102,445	$3M	0.13%	ADD 5%
84	Parkman Healthcare Partners LLC	101,237101,237	$3M	0.13%	—
85	PANAGORA ASSET MANAGEMENT INC	93,94193,941	$3M	0.12%	TRIM -29%
86	CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM	93,02993,029	$3M	0.12%	ADD 24%
87	Bridgewater Associates, LP	91,59091,590	$3M	0.11%	ADD 223%
88	LOS ANGELES CAPITAL MANAGEMENT LLC	91,40191,401	$3M	0.11%	ADD 66%
89	PRICE T ROWE ASSOCIATES INC /MD/	90,94890,948	$3M	0.11%	ADD 22%
90	AQR CAPITAL MANAGEMENT LLC	90,32290,322	$3M	0.11%	ADD 219%
91	ENVESTNET ASSET MANAGEMENT INC	86,09886,098	$3M	0.11%	ADD 38%
92	DEUTSCHE BANK AG\	85,93385,933	$3M	0.11%	ADD 15%
93	MOORE CAPITAL MANAGEMENT, LP	79,76079,760	$3M	0.10%	ADD 87%
94	Russell Investments Group, Ltd.	78,92078,920	$3M	0.10%	ADD 8%
95	CONGRESS ASSET MANAGEMENT CO	76,62776,627	$2M	0.10%	NEW
96	PERKINS CAPITAL MANAGEMENT INC	75,33375,333	$2M	0.09%	—
97	SUSQUEHANNA INTERNATIONAL GROUP, LLP	75,25875,258	$2M	0.09%	ADD 218%
98	Gotham Asset Management, LLC	72,47572,475	$2M	0.09%	ADD 4%
99	CANADA LIFE ASSURANCE Co	70,52570,525	$2M	0.09%	—
100	Empowered Funds, LLC	69,16769,167	$2M	0.09%	NEW

† a filer this large appearing as NEW usually reflects a renamed or succeeded filing entity (the old name stops filing the same quarter) — not a brand-new position.

Rows rank by the value each filer reported — 13F filers price their own positions, so a holder with fewer shares can rank just above one with more.

Holder trends· top 15 holders · shares held over 6 quarters

green added · red trimmed

Holder

2024-Q4

2025-Q1

2025-Q2

2025-Q3

2025-Q4

2026-Q1

Latest value

BlackRock, Inc.

7.5M

7.7M

13.1M

13.2M

$424M

FMR LLC

2.1M

7.1M

8.1M

6.8M

5.7M

7.8M

$250M

Artisan Partners Limited Partnership

6.6M

5.2M

6.3M

7.4M

6.3M

5.5M

$177M

STATE STREET CORP

2.9M

3.2M

4.2M

4.4M

5.0M

$162M

VANGUARD PORTFOLIO MANAGEMENT LLC

—

4.4M

$143M

VANGUARD CAPITAL MANAGEMENT LLC

—

3.5M

$114M

ARK Investment Management LLC

4.2M

3.9M

3.8M

4.5M

3.9M

3.1M

$98M

DIMENSIONAL FUND ADVISORS LP

2.9M

3.0M

3.1M

2.8M

2.9M

$93M

WELLINGTON MANAGEMENT GROUP LLP

4.1M

3.9M

3.5M

3.4M

2.0M

2.6M

$82M

FIRST TRUST ADVISORS LP

204K

1.0M

1.1M

1.3M

2.1M

$68M

GEODE CAPITAL MANAGEMENT, LLC

1.8M

1.9M

2.0M

$63M

BANK OF AMERICA CORP /DE/

743K

1.2M

1.5M

1.3M

1.4M

$47M

GW&K Investment Management, LLC

1.2M

1.3M

1.4M

1.3M

1.1M

1.4M

$46M

FULLER & THALER ASSET MANAGEMENT, INC.

—

2.8M

1.4M

$46M

ArrowMark Colorado Holdings LLC

3.1M

3.2M

2.6M

2.5M

1.6M

1.4M

$44M

Shares reported each quarter by the largest current holders. Green = added vs the prior filing, red = trimmed, grey = roughly held; a dash is a quarter with no 13F on file (a gap is skipped, not read as an exit). 13F is filed up to 45 days after quarter-end. Up to 40 quarters (10 years) available — use the selector above.

Institutional footprint by quarter

Line breaks are honest gaps — quarters with no 13F record, or a suppressed % — never interpolated. % of shares outstanding uses today's share count for every quarter (the only count stored).

Quarter	Holders	Shares held	% shares out	Value held
2026-Q1	334	88,283,972	110.6%	$2.8B
2025-Q4	328	88,188,142	—	$3.7B

Short interest has its own tab now — open SI →

Source: SEC Form 13F — every institution manager (≥ $100M in 13F securities) reporting a long US-listed position, across ~10,250 filers over 10 years of history. 13F is long-only, US-listed, filed up to 45 days after quarter-end.

Activists & large holders· Schedule 13D/G beneficial owners · VCYT

Large holders — 13G, passive (top 5 of 9 by % of class)

Holder

% of class

Type

As of

BlackRock, Inc.

15.2%

QII

2025-07-31

SEC →

FMR LLC

10.1%

QII

2026-05-29

SEC →

Artisan Partners Limited Partnership

6.9%

QII

2026-03-31

SEC →

STATE STREET CORP

6.3%

QII

2026-03-31

SEC →

VANGUARD PORTFOLIO MANAGEMENT LLC

5.67%

QII

Institutional ownership· 13F filers holding VCYT · 2026-Q1 methodology → sources →

334 holders

% of shares outstanding held

110.6%

Shares held (13F)

88,283,972

Reporting holders

334

Top holders

largest 100 of 334 · Δ vs 2025-Q4

#	Holder	Shares	Value	% of shares out	Δ QoQ
1	BlackRock, Inc.	13,172,61813.2M	$424M	16.51%	—
2	FMR LLC	7,763,8497.8M	$250M	9.73%	ADD 35%
3	Artisan Partners Limited Partnership	5,496,3705.5M	$177M	6.89%	TRIM -12%
4	STATE STREET CORP	5,043,5905.0M	$162M	6.32%	ADD 16%
5	VANGUARD PORTFOLIO MANAGEMENT LLC	4,444,7304.4M	$143M	5.57%	NEW†
6	VANGUARD CAPITAL MANAGEMENT LLC	3,532,2323.5M	$114M	4.43%	NEW†
7	ARK Investment Management LLC	3,053,8183.1M	$98M	3.83%	TRIM -21%
8	DIMENSIONAL FUND ADVISORS LP	2,881,0902.9M	$93M	3.61%	ADD 3%
9	WELLINGTON MANAGEMENT GROUP LLP	2,558,2852.6M	$82M	3.21%	ADD 26%
10	FIRST TRUST ADVISORS LP	2,117,0462.1M	$68M	2.65%	ADD 65%
11	GEODE CAPITAL MANAGEMENT, LLC	1,957,0962.0M	$63M	2.45%	ADD 3%
12	BANK OF AMERICA CORP /DE/	1,448,8971.4M	$47M	1.82%	ADD 15%
13	GW&K Investment Management, LLC	1,437,3221.4M	$46M	1.80%	ADD 26%
14	FULLER & THALER ASSET MANAGEMENT, INC.	1,414,3621.4M	$46M	1.77%	TRIM -49%
15	ArrowMark Colorado Holdings LLC	1,374,1031.4M	$44M	1.72%	TRIM -12%
16	GOLDMAN SACHS GROUP INC	1,276,2601.3M	$41M	1.60%	ADD 7%
17	Sumitomo Mitsui Trust Group, Inc.	1,055,6561.1M	$34M	1.32%	TRIM -16%
18	Amova Asset Management Americas, Inc.	1,045,0471.0M	$34M	1.31%	TRIM -17%
19	WILLIAM BLAIR INVESTMENT MANAGEMENT, LLC	989,687989,687	$32M	1.24%	ADD 25%
20	NORTHERN TRUST CORP	929,500929,500	$30M	1.16%	TRIM -6%
21	CHARLES SCHWAB INVESTMENT MANAGEMENT INC	923,573923,573	$30M	1.16%	—
22	Legal & General Group Plc	897,171897,171	$29M	1.12%	ADD 30%
23	OBERWEIS ASSET MANAGEMENT INC/	839,834839,834	$27M	1.05%	ADD 16%
24	Alyeska Investment Group, L.P.	808,570808,570	$26M	1.01%	ADD 34%
25	MORGAN STANLEY	797,802797,802	$26M	1.00%	TRIM -33%
26	Fisher Asset Management, LLC	736,270736,270	$24M	0.92%	TRIM -7%
27	DEERFIELD MANAGEMENT COMPANY, L.P.	726,000726,000	$23M	0.91%	—
28	BAMCO INC /NY/	655,522655,522	$21M	0.82%	—
29	SEGALL BRYANT & HAMILL, LLC	591,488591,488	$19M	0.74%	—
30	FRED ALGER MANAGEMENT, LLC	579,460579,460	$19M	0.73%	TRIM -9%
31	ROYAL BANK OF CANADA	573,096573,096	$18M	0.72%	ADD 66%
32	VANGUARD FIDUCIARY TRUST CO	548,167548,167	$18M	0.69%	NEW
33	Point72 Asset Management, L.P.	483,904483,904	$16M	0.61%	ADD 14%
34	PRINCIPAL FINANCIAL GROUP INC	476,686476,686	$15M	0.60%	ADD 2%
35	JPMORGAN CHASE & CO	496,219496,219	$15M	0.62%	TRIM -5%
36	Invesco Ltd.	436,587436,587	$14M	0.55%	ADD 12%
37	NEW YORK STATE COMMON RETIREMENT FUND	421,633421,633	$14M	0.53%	TRIM -28%
38	SEI INVESTMENTS CO	420,093420,093	$14M	0.53%	ADD 9%
39	Bank of New York Mellon Corp	413,416413,416	$13M	0.52%	TRIM -3%
40	BNP PARIBAS FINANCIAL MARKETS	362,836362,836	$12M	0.45%	ADD 3%
41	JUPITER ASSET MANAGEMENT LTD	355,604355,604	$11M	0.45%	—
42	RAYMOND JAMES FINANCIAL INC	370,261370,261	$11M	0.46%	TRIM -4%
43	AEGON ASSET MANAGEMENT UK Plc	326,627326,627	$11M	0.41%	TRIM -33%
44	Soleus Capital Management, L.P.	319,912319,912	$10M	0.40%	ADD 3099%
45	Balyasny Asset Management L.P.	298,955298,955	$10M	0.37%	NEW
46	Granite Investment Partners, LLC	298,589298,589	$10M	0.37%	TRIM -15%
47	Public Sector Pension Investment Board	282,835282,835	$9M	0.35%	ADD 3%
48	Champlain Investment Partners, LLC	281,923281,923	$9M	0.35%	TRIM -59%
49	Casdin Capital, LLC	276,308276,308	$9M	0.35%	—
50	Nuveen, LLC	238,439238,439	$8M	0.30%	TRIM -59%
51	Summit Creek Advisors LLC	225,894225,894	$7M	0.28%	TRIM -5%
52	RENAISSANCE TECHNOLOGIES LLC	223,500223,500	$7M	0.28%	NEW
53	NORDEA INVESTMENT MANAGEMENT AB	227,581227,581	$7M	0.29%	TRIM -4%
54	RHUMBLINE ADVISERS	213,234213,234	$7M	0.27%	—
55	FRANKLIN RESOURCES INC	211,683211,683	$7M	0.27%	—
56	UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC	205,462205,462	$7M	0.26%	ADD 4%
57	BLAIR WILLIAM & CO/IL	194,887194,887	$6M	0.24%	—
58	PRUDENTIAL FINANCIAL INC	190,910190,910	$6M	0.24%	ADD 28%
59	JANUS HENDERSON GROUP PLC	187,368187,368	$6M	0.23%	TRIM -31%
60	SILVERARC CAPITAL MANAGEMENT, LLC	179,700179,700	$6M	0.23%	TRIM -50%
61	Man Group plc	177,876177,876	$6M	0.22%	ADD 1430%
62	UBS Group AG	172,496172,496	$6M	0.22%	—
63	Palisades Investment Partners, LLC	170,603170,603	$5M	0.21%	ADD 5%
64	Allianz Asset Management GmbH	158,871158,871	$5M	0.20%	ADD 17%
65	Swiss National Bank	156,400156,400	$5M	0.20%	—
66	CITADEL ADVISORS LLC	152,941152,941	$5M	0.19%	TRIM -4%
67	NEUBERGER BERMAN GROUP LLC	150,805150,805	$5M	0.19%	ADD 2876%
68	EXCHANGE TRADED CONCEPTS, LLC	145,925145,925	$5M	0.18%	ADD 32%
69	ARROWSTREET CAPITAL, LIMITED PARTNERSHIP	141,811141,811	$5M	0.18%	NEW
70	AMERIPRISE FINANCIAL INC	135,073135,073	$4M	0.17%	TRIM -5%
71	ALLIANCEBERNSTEIN L.P.	102,643102,643	$4M	0.13%	—
72	Vestal Point Capital, LP	130,687130,687	$4M	0.16%	NEW
73	Trexquant Investment LP	130,159130,159	$4M	0.16%	ADD 1556%
74	Corient Private Wealth LLC	127,722127,722	$4M	0.16%	TRIM -51%
75	EFG International AG	125,345125,345	$4M	0.16%	TRIM -6%
76	ARDSLEY ADVISORY PARTNERS LP	125,000125,000	$4M	0.16%	—
77	IQ EQ FUND MANAGEMENT (IRELAND) Ltd	124,331124,331	$4M	0.16%	TRIM -20%
78	BROWN CAPITAL MANAGEMENT LLC	107,819107,819	$3M	0.14%	TRIM -6%
79	PINNACLE ASSOCIATES LTD	107,769107,769	$3M	0.14%	ADD 105%
80	RICE HALL JAMES & ASSOCIATES, LLC	106,665106,665	$3M	0.13%	—
81	Campbell & CO Investment Adviser LLC	106,443106,443	$3M	0.13%	ADD 581%
82	Rafferty Asset Management, LLC	104,212104,212	$3M	0.13%	ADD 21%
83	MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.	102,445102,445	$3M	0.13%	ADD 5%
84	Parkman Healthcare Partners LLC	101,237101,237	$3M	0.13%	—
85	PANAGORA ASSET MANAGEMENT INC	93,94193,941	$3M	0.12%	TRIM -29%
86	CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM	93,02993,029	$3M	0.12%	ADD 24%
87	Bridgewater Associates, LP	91,59091,590	$3M	0.11%	ADD 223%
88	LOS ANGELES CAPITAL MANAGEMENT LLC	91,40191,401	$3M	0.11%	ADD 66%
89	PRICE T ROWE ASSOCIATES INC /MD/	90,94890,948	$3M	0.11%	ADD 22%
90	AQR CAPITAL MANAGEMENT LLC	90,32290,322	$3M	0.11%	ADD 219%
91	ENVESTNET ASSET MANAGEMENT INC	86,09886,098	$3M	0.11%	ADD 38%
92	DEUTSCHE BANK AG\	85,93385,933	$3M	0.11%	ADD 15%
93	MOORE CAPITAL MANAGEMENT, LP	79,76079,760	$3M	0.10%	ADD 87%
94	Russell Investments Group, Ltd.	78,92078,920	$3M	0.10%	ADD 8%
95	CONGRESS ASSET MANAGEMENT CO	76,62776,627	$2M	0.10%	NEW
96	PERKINS CAPITAL MANAGEMENT INC	75,33375,333	$2M	0.09%	—
97	SUSQUEHANNA INTERNATIONAL GROUP, LLP	75,25875,258	$2M	0.09%	ADD 218%
98	Gotham Asset Management, LLC	72,47572,475	$2M	0.09%	ADD 4%
99	CANADA LIFE ASSURANCE Co	70,52570,525	$2M	0.09%	—
100	Empowered Funds, LLC	69,16769,167	$2M	0.09%	NEW

† a filer this large appearing as NEW usually reflects a renamed or succeeded filing entity (the old name stops filing the same quarter) — not a brand-new position.

Rows rank by the value each filer reported — 13F filers price their own positions, so a holder with fewer shares can rank just above one with more.

Holder trends· top 15 holders · shares held over 6 quarters

green added · red trimmed

Holder

2024-Q4

2025-Q1

2025-Q2

2025-Q3

2025-Q4

2026-Q1

Latest value

BlackRock, Inc.

7.5M

7.7M

13.1M

13.2M

$424M

FMR LLC

2.1M

7.1M

8.1M

6.8M

5.7M

7.8M

$250M

Artisan Partners Limited Partnership

6.6M

5.2M

6.3M

7.4M

6.3M

5.5M

$177M

STATE STREET CORP

2.9M

3.2M

4.2M

4.4M

5.0M

$162M

VANGUARD PORTFOLIO MANAGEMENT LLC

—

4.4M

$143M

VANGUARD CAPITAL MANAGEMENT LLC

—

3.5M

$114M

ARK Investment Management LLC

4.2M

3.9M

3.8M

4.5M

3.9M

3.1M

$98M

DIMENSIONAL FUND ADVISORS LP

2.9M

3.0M

3.1M

2.8M

2.9M

$93M

WELLINGTON MANAGEMENT GROUP LLP

4.1M

3.9M

3.5M

3.4M

2.0M

2.6M

$82M

FIRST TRUST ADVISORS LP

204K

1.0M

1.1M

1.3M

2.1M

$68M

GEODE CAPITAL MANAGEMENT, LLC

1.8M

1.9M

2.0M

$63M

BANK OF AMERICA CORP /DE/

743K

1.2M

1.5M

1.3M

1.4M

$47M

GW&K Investment Management, LLC

1.2M

1.3M

1.4M

1.3M

1.1M

1.4M

$46M

FULLER & THALER ASSET MANAGEMENT, INC.

—

2.8M

1.4M

$46M

ArrowMark Colorado Holdings LLC

3.1M

3.2M

2.6M

2.5M

1.6M

1.4M

$44M

Institutional footprint by quarter

Line breaks are honest gaps — quarters with no 13F record, or a suppressed % — never interpolated. % of shares outstanding uses today's share count for every quarter (the only count stored).

Quarter	Holders	Shares held	% shares out	Value held
2026-Q1	334	88,283,972	110.6%	$2.8B
2025-Q4	328	88,188,142	—	$3.7B

Short interest has its own tab now — open SI →

Activists & large holders· Schedule 13D/G beneficial owners · VCYT

Large holders — 13G, passive (top 5 of 9 by % of class)

Holder

% of class

Type

As of

BlackRock, Inc.

15.2%

QII

2025-07-31

SEC →

FMR LLC

10.1%

QII

2026-05-29

SEC →

Artisan Partners Limited Partnership

6.9%

QII

2026-03-31

SEC →

STATE STREET CORP

6.3%

QII

2026-03-31

SEC →

VANGUARD PORTFOLIO MANAGEMENT LLC

5.67%

QII

VCYT US Equity☆

Institutional ownership· 13F filers holding VCYT · 2026-Q1 methodology → sources →

Top holders

Holder trends· top 15 holders · shares held over 6 quarters

Institutional footprint by quarter

Activists & large holders· Schedule 13D/G beneficial owners · VCYT

Institutional ownership· 13F filers holding VCYT · 2026-Q1 methodology → sources →

Top holders

Holder trends· top 15 holders · shares held over 6 quarters

Institutional footprint by quarter

Activists & large holders· Schedule 13D/G beneficial owners · VCYT

VCYT US Equity