ATYR US Equity

aTYR PHARMA INC

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Health Care · Biological Products, (No Diagnostic Substances) · CIK 1339970 · FYE 1231

$0.48

-0.01 (-2.23%)

USD · as of 2026-07-29 · marketstack

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Institutional ownership· 13F filers holding ATYR · 2026-Q1 methodology → sources →

111 holders

% of shares outstanding held

57.2%

Shares held (13F)

56,056,214

Reporting holders

111

Top holders

largest 100 of 111 · Δ vs 2025-Q4

#	Holder	Shares	Value	% of shares out	Δ QoQ
1	FMR LLC	14,706,84314.7M	$11M	15.00%	—
2	FEDERATED HERMES, INC.	10,509,67810.5M	$8M	10.72%	—
3	BlackRock, Inc.	6,443,9046.4M	$5M	6.57%	—
4	UBS Group AG	3,867,2133.9M	$3M	3.94%	ADD 138%
5	VANGUARD CAPITAL MANAGEMENT LLC	3,812,1323.8M	$3M	3.89%	NEW†
6	GEODE CAPITAL MANAGEMENT, LLC	2,385,4852.4M	$2M	2.43%	ADD 4%
7	STATE STREET CORP	1,754,3991.8M	$1M	1.79%	—
8	GSA CAPITAL PARTNERS LLP	1,684,5861.7M	$1M	1.72%	TRIM -36%
9	MILLENNIUM MANAGEMENT LLC	1,641,2461.6M	$1M	1.67%	NEW
10	MORGAN STANLEY	1,225,4671.2M	$955,865	1.25%	TRIM -13%
11	TWO SIGMA INVESTMENTS, LP	1,144,5651.1M	$892,761	1.17%	ADD 152%
12	SUSQUEHANNA INTERNATIONAL GROUP, LLP	866,744866,744	$676,060	0.88%	ADD 13%
13	NORTHERN TRUST CORP	711,105711,105	$554,663	0.73%	—
14	VANGUARD FIDUCIARY TRUST CO	651,832651,832	$508,429	0.66%	NEW
15	VANGUARD PORTFOLIO MANAGEMENT LLC	569,227569,227	$443,997	0.58%	NEW
16	AQR CAPITAL MANAGEMENT LLC	496,018496,018	$386,894	0.51%	TRIM -36%
17	BANK OF AMERICA CORP /DE/	303,559303,559	$236,776	0.31%	ADD 140%
18	GOLDMAN SACHS GROUP INC	283,236283,236	$220,924	0.29%	TRIM -18%
19	Nuveen, LLC	229,284229,284	$178,842	0.23%	—
20	CHARLES SCHWAB INVESTMENT MANAGEMENT INC	210,737210,737	$164,375	0.21%	TRIM -5%
21	Bank of New York Mellon Corp	193,531193,531	$150,954	0.20%	—
22	GROUP ONE TRADING LLC	184,014184,014	$143,531	0.19%	TRIM -4%
23	MARSHALL WACE, LLP	138,118138,118	$107,733	0.14%	NEW
24	XTX Topco Ltd	111,423111,423	$86,910	0.11%	ADD 40%
25	DEUTSCHE BANK AG\	103,326103,326	$80,595	0.11%	TRIM -14%
26	RHUMBLINE ADVISERS	103,146103,146	$80,445	0.11%	ADD 3%
27	HOHIMER WEALTH MANAGEMENT, LLC	100,000100,000	$78,000	0.10%	NEW
28	DIMENSIONAL FUND ADVISORS LP	91,87191,871	$71,479	0.09%	TRIM -6%
29	VICTORY CAPITAL MANAGEMENT INC	88,43688,436	$68,980	0.09%	TRIM -51%
30	ALLIANCEBERNSTEIN L.P.	64,48064,480	$50,494	0.07%	—
31	KINGSWOOD WEALTH ADVISORS, LLC	52,00052,000	$40,560	0.05%	—
32	ADAR1 Capital Management, LLC	50,68550,685	$39,534	0.05%	—
33	Redhawk Wealth Advisors, Inc.	49,15049,150	$38,337	0.05%	NEW
34	JANE STREET GROUP, LLC	48,50248,502	$37,832	0.05%	TRIM -74%
35	CITADEL ADVISORS LLC	47,37247,372	$36,950	0.05%	NEW
36	WELLS FARGO & COMPANY/MN	46,54046,540	$36,301	0.05%	TRIM -60%
37	Invesco Ltd.	46,35446,354	$36,156	0.05%	—
38	PRICE T ROWE ASSOCIATES INC /MD/	46,12646,126	$36,000	0.05%	—
39	MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.	46,07346,073	$35,937	0.05%	ADD 10%
40	Belpointe Asset Management LLC	45,73845,738	$35,676	0.05%	ADD 338%
41	BARCLAYS PLC	45,12345,123	$35,196	0.05%	TRIM -74%
42	RENAISSANCE TECHNOLOGIES LLC	40,52240,522	$31,607	0.04%	NEW
43	Virtu Financial LLC	40,04840,048	$31,000	0.04%	TRIM -49%
44	JPMORGAN CHASE & CO	41,98741,987	$30,689	0.04%	TRIM -62%
45	CWM, LLC	36,01036,010	$28,088	0.04%	ADD 351%
46	LPL Financial LLC	35,54935,549	$27,728	0.04%	ADD 18%
47	SEI INVESTMENTS CO	33,82933,829	$26,387	0.03%	—
48	Vanguard Global Advisers, LLC	33,41433,414	$26,063	0.03%	NEW
49	Rockefeller Capital Management L.P.	28,11828,118	$21,932	0.03%	—
50	NEW YORK STATE COMMON RETIREMENT FUND	27,50027,500	$21,450	0.03%	—
51	INTECH INVESTMENT MANAGEMENT LLC	26,18726,187	$20,426	0.03%	TRIM -18%
52	Police & Firemen's Retirement System of New Jersey	26,08126,081	$20,343	0.03%	—
53	BNP PARIBAS FINANCIAL MARKETS	25,27825,278	$19,717	0.03%	TRIM -5%
54	NEUBERGER BERMAN GROUP LLC	21,74321,743	$16,959	0.02%	ADD 6%
55	RBF Capital, LLC	20,00020,000	$15,600	0.02%	TRIM -50%
56	Hobbs Group Advisors, LLC	19,41919,419	$15,147	0.02%	NEW
57	HSBC HOLDINGS PLC	18,21418,214	$13,870	0.02%	TRIM -44%
58	Tower Research Capital LLC (TRC)	17,42617,426	$13,592	0.02%	ADD 27%
59	WEALTH ENHANCEMENT ADVISORY SERVICES, LLC	15,57815,578	$13,193	0.02%	NEW
60	Vanguard Personalized Indexing Management, LLC	16,66916,669	$13,002	0.02%	NEW
61	TrinityBridge Ltd	16,20016,200	$13,000	0.02%	NEW
62	TRUST CO OF TOLEDO NA /OH/	16,64016,640	$12,979	0.02%	NEW
63	RAYMOND JAMES FINANCIAL INC	16,60116,601	$12,949	0.02%	ADD 4%
64	Steward Partners Investment Advisory, LLC	16,60016,600	$12,948	0.02%	NEW
65	CAPTRUST FINANCIAL ADVISORS	15,98215,982	$12,462	0.02%	NEW
66	MERCER GLOBAL ADVISORS INC /ADV	15,20515,205	$11,860	0.02%	NEW
67	PRUDENTIAL FINANCIAL INC	14,66014,660	$11,435	0.01%	NEW
68	ProShare Advisors LLC	13,90513,905	$10,845	0.01%	TRIM -21%
69	MAI Capital Management	13,50413,504	$10,534	0.01%	TRIM -11%
70	Crossmark Global Holdings, Inc.	12,66612,666	$9,879	0.01%	—
71	Pasadena Private Wealth, LLC	12,22812,228	$9,538	0.01%	NEW
72	Apollon Wealth Management, LLC	11,90011,900	$9,282	0.01%	TRIM -8%
73	ROYAL BANK OF CANADA	11,14611,146	$9,000	0.01%	ADD 112%
74	Russell Investments Group, Ltd.	11,36111,361	$8,861	0.01%	ADD 58%
75	Gladstone Institutional Advisory LLC	11,30011,300	$8,814	0.01%	—
76	MANUFACTURERS LIFE INSURANCE COMPANY, THE	10,55210,552	$8,231	0.01%	NEW
77	Creative Planning	10,50810,508	$8,196	0.01%	NEW
78	VOYA INVESTMENT MANAGEMENT LLC	10,33810,338	$8,064	0.01%	TRIM -38%
79	Balyasny Asset Management L.P.	10,33010,330	$8,057	0.01%	NEW
80	Ameritas Investment Partners, Inc.	8,8818,881	$6,927	0.01%	—
81	PNC FINANCIAL SERVICES GROUP, INC.	8,1968,196	$6,393	0.01%	ADD 1615%
82	Legal & General Group Plc	7,9207,920	$6,178	0.01%	TRIM -17%
83	GAMMA Investing LLC	7,7397,739	$6,036	0.01%	NEW
84	My Portfolio Guide, LLC	7,0007,000	$5,460	0.01%	NEW
85	IFP Advisors, Inc	6,0016,001	$4,681	0.01%	NEW
86	CITIGROUP INC	5,8915,891	$4,595	0.01%	ADD 92%
87	CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM	5,1905,190	$4,048	0.01%	TRIM -12%
88	OSAIC HOLDINGS, INC.	5,0005,000	$3,900	0.01%	NEW
89	Ares Financial Consulting, LLC	2,0002,000	$1,702	0.00%	NEW
90	NFSG Corp	2,0792,079	$1,622	0.00%	TRIM -42%
91	UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC	2,0112,011	$1,569	0.00%	NEW
92	Essential Partners LLC	1,6181,618	$1,262	0.00%	ADD 825%
93	Vision Retirement, LLC	1,5001,500	$1,170	0.00%	—
94	JONES FINANCIAL COMPANIES LLLP	1,5001,500	$1,128	0.00%	—
95	Farther Finance Advisors, LLC	1,3201,320	$1,123	0.00%	TRIM -41%
96	TOTH FINANCIAL ADVISORY CORP	1,4001,400	$1,092	0.00%	NEW
97	SIMPLEX TRADING, LLC	1,3351,335	$1,041	0.00%	TRIM -95%
98	FNY Investment Advisers, LLC	1,5001,500	$1,000	0.00%	TRIM -69%
99	NISA INVESTMENT ADVISORS, LLC	974974	$760	0.00%	NEW
100	Sterling Capital Management LLC	742742	$579	0.00%	TRIM -21%

† a filer this large appearing as NEW usually reflects a renamed or succeeded filing entity (the old name stops filing the same quarter) — not a brand-new position.

Rows rank by the value each filer reported — 13F filers price their own positions, so a holder with fewer shares can rank just above one with more.

Holder trends· top 15 holders · shares held over 6 quarters

green added · red trimmed

Holder

2024-Q4

2025-Q1

2025-Q2

2025-Q3

2025-Q4

2026-Q1

Latest value

FMR LLC

12.1M

12.9M

13.4M

14.7M

$11M

FEDERATED HERMES, INC.

14.7M

10.5M

$8M

BlackRock, Inc.

1.5M

1.6M

5.8M

6.5M

6.4M

$5M

UBS Group AG

230K

1.6M

1.7M

4.5M

1.6M

3.9M

$3M

VANGUARD CAPITAL MANAGEMENT LLC

—

3.8M

$3M

GEODE CAPITAL MANAGEMENT, LLC

891K

928K

2.1M

2.2M

2.3M

2.4M

$2M

STATE STREET CORP

301K

367K

1.2M

4.2M

1.7M

1.8M

$1M

GSA CAPITAL PARTNERS LLP

274K

332K

496K

707K

2.6M

1.7M

$1M

MILLENNIUM MANAGEMENT LLC

368K

1.6M

1.9M

—

1.6M

$1M

MORGAN STANLEY

41K

263K

260K

710K

1.4M

1.2M

$955,865

TWO SIGMA INVESTMENTS, LP

22K

31K

211K

454K

1.1M

$892,761

SUSQUEHANNA INTERNATIONAL GROUP, LLP

788K

1.7M

254K

127K

765K

867K

$676,060

NORTHERN TRUST CORP

183K

187K

681K

674K

702K

711K

$554,663

VANGUARD FIDUCIARY TRUST CO

—

652K

$508,429

VANGUARD PORTFOLIO MANAGEMENT LLC

—

569K

$443,997

Shares reported each quarter by the largest current holders. Green = added vs the prior filing, red = trimmed, grey = roughly held; a dash is a quarter with no 13F on file (a gap is skipped, not read as an exit). 13F is filed up to 45 days after quarter-end. Up to 40 quarters (10 years) available — use the selector above.

Institutional footprint by quarter

Line breaks are honest gaps — quarters with no 13F record, or a suppressed % — never interpolated. % of shares outstanding uses today's share count for every quarter (the only count stored).

Quarter	Holders	Shares held	% shares out	Value held
2026-Q1	111	56,056,214	57.2%	$44M
2025-Q4	84	54,865,270	56.0%	$56M
2025-Q3	103	62,045,169	63.3%	$52M
2025-Q2	110	66,458,240	67.8%

Short interest has its own tab now — open SI →

Source: SEC Form 13F — every institution manager (≥ $100M in 13F securities) reporting a long US-listed position, across ~10,250 filers over 10 years of history. 13F is long-only, US-listed, filed up to 45 days after quarter-end.

Activists & large holders· Schedule 13D/G beneficial owners · ATYR

Large holders — 13G, passive (top 5 by % of class)

Holder	% of class	Type	As of
FEDERATED HERMES, INC.	8.68%	QII	2026-06-30	SEC →
FMR LLC	6.1%	QII	2026-05-29	SEC →
Point72 Asset Management, L.P.	3.2%	Passive	2025-03-31	SEC →
BlackRock, Inc.	1.8%	QII	2026-06-30	SEC →
VANGUARD GROUP INC	0%	QII	2026-03-13

Institutional ownership· 13F filers holding ATYR · 2026-Q1 methodology → sources →

111 holders

% of shares outstanding held

57.2%

Shares held (13F)

56,056,214

Reporting holders

111

Top holders

largest 100 of 111 · Δ vs 2025-Q4

#	Holder	Shares	Value	% of shares out	Δ QoQ
1	FMR LLC	14,706,84314.7M	$11M	15.00%	—
2	FEDERATED HERMES, INC.	10,509,67810.5M	$8M	10.72%	—
3	BlackRock, Inc.	6,443,9046.4M	$5M	6.57%	—
4	UBS Group AG	3,867,2133.9M	$3M	3.94%	ADD 138%
5	VANGUARD CAPITAL MANAGEMENT LLC	3,812,1323.8M	$3M	3.89%	NEW†
6	GEODE CAPITAL MANAGEMENT, LLC	2,385,4852.4M	$2M	2.43%	ADD 4%
7	STATE STREET CORP	1,754,3991.8M	$1M	1.79%	—
8	GSA CAPITAL PARTNERS LLP	1,684,5861.7M	$1M	1.72%	TRIM -36%
9	MILLENNIUM MANAGEMENT LLC	1,641,2461.6M	$1M	1.67%	NEW
10	MORGAN STANLEY	1,225,4671.2M	$955,865	1.25%	TRIM -13%
11	TWO SIGMA INVESTMENTS, LP	1,144,5651.1M	$892,761	1.17%	ADD 152%
12	SUSQUEHANNA INTERNATIONAL GROUP, LLP	866,744866,744	$676,060	0.88%	ADD 13%
13	NORTHERN TRUST CORP	711,105711,105	$554,663	0.73%	—
14	VANGUARD FIDUCIARY TRUST CO	651,832651,832	$508,429	0.66%	NEW
15	VANGUARD PORTFOLIO MANAGEMENT LLC	569,227569,227	$443,997	0.58%	NEW
16	AQR CAPITAL MANAGEMENT LLC	496,018496,018	$386,894	0.51%	TRIM -36%
17	BANK OF AMERICA CORP /DE/	303,559303,559	$236,776	0.31%	ADD 140%
18	GOLDMAN SACHS GROUP INC	283,236283,236	$220,924	0.29%	TRIM -18%
19	Nuveen, LLC	229,284229,284	$178,842	0.23%	—
20	CHARLES SCHWAB INVESTMENT MANAGEMENT INC	210,737210,737	$164,375	0.21%	TRIM -5%
21	Bank of New York Mellon Corp	193,531193,531	$150,954	0.20%	—
22	GROUP ONE TRADING LLC	184,014184,014	$143,531	0.19%	TRIM -4%
23	MARSHALL WACE, LLP	138,118138,118	$107,733	0.14%	NEW
24	XTX Topco Ltd	111,423111,423	$86,910	0.11%	ADD 40%
25	DEUTSCHE BANK AG\	103,326103,326	$80,595	0.11%	TRIM -14%
26	RHUMBLINE ADVISERS	103,146103,146	$80,445	0.11%	ADD 3%
27	HOHIMER WEALTH MANAGEMENT, LLC	100,000100,000	$78,000	0.10%	NEW
28	DIMENSIONAL FUND ADVISORS LP	91,87191,871	$71,479	0.09%	TRIM -6%
29	VICTORY CAPITAL MANAGEMENT INC	88,43688,436	$68,980	0.09%	TRIM -51%
30	ALLIANCEBERNSTEIN L.P.	64,48064,480	$50,494	0.07%	—
31	KINGSWOOD WEALTH ADVISORS, LLC	52,00052,000	$40,560	0.05%	—
32	ADAR1 Capital Management, LLC	50,68550,685	$39,534	0.05%	—
33	Redhawk Wealth Advisors, Inc.	49,15049,150	$38,337	0.05%	NEW
34	JANE STREET GROUP, LLC	48,50248,502	$37,832	0.05%	TRIM -74%
35	CITADEL ADVISORS LLC	47,37247,372	$36,950	0.05%	NEW
36	WELLS FARGO & COMPANY/MN	46,54046,540	$36,301	0.05%	TRIM -60%
37	Invesco Ltd.	46,35446,354	$36,156	0.05%	—
38	PRICE T ROWE ASSOCIATES INC /MD/	46,12646,126	$36,000	0.05%	—
39	MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.	46,07346,073	$35,937	0.05%	ADD 10%
40	Belpointe Asset Management LLC	45,73845,738	$35,676	0.05%	ADD 338%
41	BARCLAYS PLC	45,12345,123	$35,196	0.05%	TRIM -74%
42	RENAISSANCE TECHNOLOGIES LLC	40,52240,522	$31,607	0.04%	NEW
43	Virtu Financial LLC	40,04840,048	$31,000	0.04%	TRIM -49%
44	JPMORGAN CHASE & CO	41,98741,987	$30,689	0.04%	TRIM -62%
45	CWM, LLC	36,01036,010	$28,088	0.04%	ADD 351%
46	LPL Financial LLC	35,54935,549	$27,728	0.04%	ADD 18%
47	SEI INVESTMENTS CO	33,82933,829	$26,387	0.03%	—
48	Vanguard Global Advisers, LLC	33,41433,414	$26,063	0.03%	NEW
49	Rockefeller Capital Management L.P.	28,11828,118	$21,932	0.03%	—
50	NEW YORK STATE COMMON RETIREMENT FUND	27,50027,500	$21,450	0.03%	—
51	INTECH INVESTMENT MANAGEMENT LLC	26,18726,187	$20,426	0.03%	TRIM -18%
52	Police & Firemen's Retirement System of New Jersey	26,08126,081	$20,343	0.03%	—
53	BNP PARIBAS FINANCIAL MARKETS	25,27825,278	$19,717	0.03%	TRIM -5%
54	NEUBERGER BERMAN GROUP LLC	21,74321,743	$16,959	0.02%	ADD 6%
55	RBF Capital, LLC	20,00020,000	$15,600	0.02%	TRIM -50%
56	Hobbs Group Advisors, LLC	19,41919,419	$15,147	0.02%	NEW
57	HSBC HOLDINGS PLC	18,21418,214	$13,870	0.02%	TRIM -44%
58	Tower Research Capital LLC (TRC)	17,42617,426	$13,592	0.02%	ADD 27%
59	WEALTH ENHANCEMENT ADVISORY SERVICES, LLC	15,57815,578	$13,193	0.02%	NEW
60	Vanguard Personalized Indexing Management, LLC	16,66916,669	$13,002	0.02%	NEW
61	TrinityBridge Ltd	16,20016,200	$13,000	0.02%	NEW
62	TRUST CO OF TOLEDO NA /OH/	16,64016,640	$12,979	0.02%	NEW
63	RAYMOND JAMES FINANCIAL INC	16,60116,601	$12,949	0.02%	ADD 4%
64	Steward Partners Investment Advisory, LLC	16,60016,600	$12,948	0.02%	NEW
65	CAPTRUST FINANCIAL ADVISORS	15,98215,982	$12,462	0.02%	NEW
66	MERCER GLOBAL ADVISORS INC /ADV	15,20515,205	$11,860	0.02%	NEW
67	PRUDENTIAL FINANCIAL INC	14,66014,660	$11,435	0.01%	NEW
68	ProShare Advisors LLC	13,90513,905	$10,845	0.01%	TRIM -21%
69	MAI Capital Management	13,50413,504	$10,534	0.01%	TRIM -11%
70	Crossmark Global Holdings, Inc.	12,66612,666	$9,879	0.01%	—
71	Pasadena Private Wealth, LLC	12,22812,228	$9,538	0.01%	NEW
72	Apollon Wealth Management, LLC	11,90011,900	$9,282	0.01%	TRIM -8%
73	ROYAL BANK OF CANADA	11,14611,146	$9,000	0.01%	ADD 112%
74	Russell Investments Group, Ltd.	11,36111,361	$8,861	0.01%	ADD 58%
75	Gladstone Institutional Advisory LLC	11,30011,300	$8,814	0.01%	—
76	MANUFACTURERS LIFE INSURANCE COMPANY, THE	10,55210,552	$8,231	0.01%	NEW
77	Creative Planning	10,50810,508	$8,196	0.01%	NEW
78	VOYA INVESTMENT MANAGEMENT LLC	10,33810,338	$8,064	0.01%	TRIM -38%
79	Balyasny Asset Management L.P.	10,33010,330	$8,057	0.01%	NEW
80	Ameritas Investment Partners, Inc.	8,8818,881	$6,927	0.01%	—
81	PNC FINANCIAL SERVICES GROUP, INC.	8,1968,196	$6,393	0.01%	ADD 1615%
82	Legal & General Group Plc	7,9207,920	$6,178	0.01%	TRIM -17%
83	GAMMA Investing LLC	7,7397,739	$6,036	0.01%	NEW
84	My Portfolio Guide, LLC	7,0007,000	$5,460	0.01%	NEW
85	IFP Advisors, Inc	6,0016,001	$4,681	0.01%	NEW
86	CITIGROUP INC	5,8915,891	$4,595	0.01%	ADD 92%
87	CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM	5,1905,190	$4,048	0.01%	TRIM -12%
88	OSAIC HOLDINGS, INC.	5,0005,000	$3,900	0.01%	NEW
89	Ares Financial Consulting, LLC	2,0002,000	$1,702	0.00%	NEW
90	NFSG Corp	2,0792,079	$1,622	0.00%	TRIM -42%
91	UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC	2,0112,011	$1,569	0.00%	NEW
92	Essential Partners LLC	1,6181,618	$1,262	0.00%	ADD 825%
93	Vision Retirement, LLC	1,5001,500	$1,170	0.00%	—
94	JONES FINANCIAL COMPANIES LLLP	1,5001,500	$1,128	0.00%	—
95	Farther Finance Advisors, LLC	1,3201,320	$1,123	0.00%	TRIM -41%
96	TOTH FINANCIAL ADVISORY CORP	1,4001,400	$1,092	0.00%	NEW
97	SIMPLEX TRADING, LLC	1,3351,335	$1,041	0.00%	TRIM -95%
98	FNY Investment Advisers, LLC	1,5001,500	$1,000	0.00%	TRIM -69%
99	NISA INVESTMENT ADVISORS, LLC	974974	$760	0.00%	NEW
100	Sterling Capital Management LLC	742742	$579	0.00%	TRIM -21%

† a filer this large appearing as NEW usually reflects a renamed or succeeded filing entity (the old name stops filing the same quarter) — not a brand-new position.

Rows rank by the value each filer reported — 13F filers price their own positions, so a holder with fewer shares can rank just above one with more.

Holder trends· top 15 holders · shares held over 6 quarters

green added · red trimmed

Holder

2024-Q4

2025-Q1

2025-Q2

2025-Q3

2025-Q4

2026-Q1

Latest value

FMR LLC

12.1M

12.9M

13.4M

14.7M

$11M

FEDERATED HERMES, INC.

14.7M

10.5M

$8M

BlackRock, Inc.

1.5M

1.6M

5.8M

6.5M

6.4M

$5M

UBS Group AG

230K

1.6M

1.7M

4.5M

1.6M

3.9M

$3M

VANGUARD CAPITAL MANAGEMENT LLC

—

3.8M

$3M

GEODE CAPITAL MANAGEMENT, LLC

891K

928K

2.1M

2.2M

2.3M

2.4M

$2M

STATE STREET CORP

301K

367K

1.2M

4.2M

1.7M

1.8M

$1M

GSA CAPITAL PARTNERS LLP

274K

332K

496K

707K

2.6M

1.7M

$1M

MILLENNIUM MANAGEMENT LLC

368K

1.6M

1.9M

—

1.6M

$1M

MORGAN STANLEY

41K

263K

260K

710K

1.4M

1.2M

$955,865

TWO SIGMA INVESTMENTS, LP

22K

31K

211K

454K

1.1M

$892,761

SUSQUEHANNA INTERNATIONAL GROUP, LLP

788K

1.7M

254K

127K

765K

867K

$676,060

NORTHERN TRUST CORP

183K

187K

681K

674K

702K

711K

$554,663

VANGUARD FIDUCIARY TRUST CO

—

652K

$508,429

VANGUARD PORTFOLIO MANAGEMENT LLC

—

569K

$443,997

Institutional footprint by quarter

Line breaks are honest gaps — quarters with no 13F record, or a suppressed % — never interpolated. % of shares outstanding uses today's share count for every quarter (the only count stored).

Quarter	Holders	Shares held	% shares out	Value held
2026-Q1	111	56,056,214	57.2%	$44M
2025-Q4	84	54,865,270	56.0%	$56M
2025-Q3	103	62,045,169	63.3%	$52M
2025-Q2	110	66,458,240	67.8%

Short interest has its own tab now — open SI →

Activists & large holders· Schedule 13D/G beneficial owners · ATYR

Large holders — 13G, passive (top 5 by % of class)

Holder	% of class	Type	As of
FEDERATED HERMES, INC.	8.68%	QII	2026-06-30	SEC →
FMR LLC	6.1%	QII	2026-05-29	SEC →
Point72 Asset Management, L.P.	3.2%	Passive	2025-03-31	SEC →
BlackRock, Inc.	1.8%	QII	2026-06-30	SEC →
VANGUARD GROUP INC	0%	QII	2026-03-13

ATYR US Equity☆

Institutional ownership· 13F filers holding ATYR · 2026-Q1 methodology → sources →

Top holders

Holder trends· top 15 holders · shares held over 6 quarters

Institutional footprint by quarter

Activists & large holders· Schedule 13D/G beneficial owners · ATYR

Institutional ownership· 13F filers holding ATYR · 2026-Q1 methodology → sources →

Top holders

Holder trends· top 15 holders · shares held over 6 quarters

Institutional footprint by quarter

Activists & large holders· Schedule 13D/G beneficial owners · ATYR

ATYR US Equity