APPN US Equity

Name: SPZCO
Price: 29.99 USD

APPIAN CORP

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Information Technology · Services-Prepackaged Software · CIK 1441683 · FYE 1231

$28.74

+1.39 (+5.08%)

USD · as of 2026-07-29 · marketstack

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Institutional ownership· 13F filers holding APPN · 2026-Q1 methodology → sources →

206 holders

% of shares outstanding held

52.1%

Shares held (13F)

38,913,967

Reporting holders

206

Top holders

largest 100 of 206 · Δ vs 2025-Q4

#	Holder	Shares	Value	% of shares out	Δ QoQ
1	VANGUARD PORTFOLIO MANAGEMENT LLC	4,161,5594.2M	$100M	5.57%	NEW†
2	RPD Fund Management LLC	3,919,3563.9M	$94M	5.25%	NEW†
3	Fivespan Partners, LP	3,510,9073.5M	$85M	4.70%	ADD 155%
4	BlackRock, Inc.	3,226,0873.2M	$78M	4.32%	—
5	Lead Edge Capital Management, LLC	1,905,6341.9M	$46M	2.55%	ADD 18%
6	VANGUARD CAPITAL MANAGEMENT LLC	1,678,2801.7M	$40M	2.25%	NEW†
7	MORGAN STANLEY	1,545,8631.5M	$37M	2.07%	ADD 9%
8	ARROWSTREET CAPITAL, LIMITED PARTNERSHIP	1,519,6531.5M	$37M	2.04%	ADD 16%
9	FIRST TRUST ADVISORS LP	1,355,9611.4M	$33M	1.82%	ADD 17%
10	STATE STREET CORP	1,023,4421.0M	$25M	1.37%	ADD 4%
11	GEODE CAPITAL MANAGEMENT, LLC	985,499985,499	$24M	1.32%	—
12	AQR CAPITAL MANAGEMENT LLC	957,620957,620	$23M	1.28%	ADD 43%
13	BAILLIE GIFFORD & CO	905,908905,908	$22M	1.21%	TRIM -4%
14	Mudita Advisors LLP	839,400839,400	$20M	1.12%	NEW
15	JACOBS LEVY EQUITY MANAGEMENT, INC	603,430603,430	$15M	0.81%	TRIM -3%
16	ACADIAN ASSET MANAGEMENT LLC	520,419520,419	$13M	0.70%	TRIM -15%
17	CITADEL ADVISORS LLC	518,009518,009	$12M	0.69%	TRIM -6%
18	D. E. Shaw & Co., Inc.	447,788447,788	$11M	0.60%	TRIM -60%
19	Nuveen, LLC	414,956414,956	$10M	0.56%	TRIM -40%
20	Squarepoint Ops LLC	390,929390,929	$9M	0.52%	ADD 32%
21	PRELUDE CAPITAL MANAGEMENT, LLC	390,321390,321	$9M	0.52%	NEW
22	FMR LLC	380,697380,697	$9M	0.51%	ADD 18%
23	CHARLES SCHWAB INVESTMENT MANAGEMENT INC	370,662370,662	$9M	0.50%	ADD 6%
24	MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.	353,022353,022	$9M	0.47%	TRIM -33%
25	NORTHERN TRUST CORP	344,336344,336	$8M	0.46%	ADD 2%
26	Trexquant Investment LP	305,943305,943	$7M	0.41%	ADD 97%
27	National Philanthropic Trust	299,532299,532	$7M	0.40%	—
28	Qube Research & Technologies Ltd	292,737292,737	$7M	0.39%	TRIM -39%
29	GOLDMAN SACHS GROUP INC	285,265285,265	$7M	0.38%	TRIM -30%
30	UBS Group AG	284,284284,284	$7M	0.38%	ADD 205%
31	VANGUARD FIDUCIARY TRUST CO	277,406277,406	$7M	0.37%	NEW
32	Connor, Clark & Lunn Investment Management Ltd.	208,561208,561	$5M	0.28%	TRIM -43%
33	JANE STREET GROUP, LLC	194,452194,452	$5M	0.26%	TRIM -23%
34	HRT FINANCIAL LP	189,984189,984	$5M	0.25%	ADD 79%
35	ALGERT GLOBAL LLC	179,554179,554	$4M	0.24%	ADD 23%
36	CANADA PENSION PLAN INVESTMENT BOARD	179,500179,500	$4M	0.24%	TRIM -9%
37	BANK OF AMERICA CORP /DE/	176,566176,566	$4M	0.24%	ADD 8%
38	Tejara Capital Ltd	173,300173,300	$4M	0.23%	NEW
39	Harber Asset Management LLC	147,426147,426	$4M	0.20%	TRIM -22%
40	Point72 Asset Management, L.P.	147,228147,228	$4M	0.20%	TRIM -36%
41	ExodusPoint Capital Management, LP	146,023146,023	$4M	0.20%	NEW
42	JPMORGAN CHASE & CO	145,224145,224	$3M	0.19%	ADD 30%
43	Engineers Gate Manager LP	130,979130,979	$3M	0.18%	TRIM -15%
44	BNP PARIBAS FINANCIAL MARKETS	127,268127,268	$3M	0.17%	TRIM -48%
45	Russell Investments Group, Ltd.	123,051123,051	$3M	0.16%	TRIM -49%
46	Bank of New York Mellon Corp	120,429120,429	$3M	0.16%	TRIM -2%
47	TWO SIGMA INVESTMENTS, LP	119,066119,066	$3M	0.16%	TRIM -63%
48	Gotham Asset Management, LLC	95,64895,648	$2M	0.13%	ADD 99%
49	UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC	86,00086,000	$2M	0.12%	ADD 2%
50	SEI INVESTMENTS CO	85,09985,099	$2M	0.11%	ADD 8%
51	LOS ANGELES CAPITAL MANAGEMENT LLC	80,62480,624	$2M	0.11%	ADD 107%
52	Swiss National Bank	76,30076,300	$2M	0.10%	—
53	BOOTHBAY FUND MANAGEMENT, LLC	75,16075,160	$2M	0.10%	NEW
54	BARCLAYS PLC	74,74874,748	$2M	0.10%	TRIM -44%
55	ALLIANCEBERNSTEIN L.P.	44,56144,561	$2M	0.06%	ADD 5%
56	IEQ CAPITAL, LLC	63,24863,248	$2M	0.08%	TRIM -24%
57	Brevan Howard Capital Management LP	58,89858,898	$1M	0.08%	ADD 495%
58	CWM, LLC	53,82053,820	$1M	0.07%	ADD 65%
59	RHUMBLINE ADVISERS	51,62751,627	$1M	0.07%	TRIM -25%
60	PRICE T ROWE ASSOCIATES INC /MD/	49,22649,226	$1M	0.07%	ADD 18%
61	Magnetar Financial LLC	48,07148,071	$1M	0.06%	ADD 81%
62	FRANKLIN RESOURCES INC	47,23847,238	$1M	0.06%	ADD 186%
63	PDT Partners, LLC	47,08947,089	$1M	0.06%	TRIM -58%
64	Invesco Ltd.	46,98446,984	$1M	0.06%	—
65	CITIGROUP INC	45,91945,919	$1M	0.06%	TRIM -79%
66	Mariner, LLC	42,87442,874	$1M	0.06%	ADD 78%
67	Amundi	42,80442,804	$1M	0.06%	TRIM -67%
68	Man Group plc	42,44042,440	$1M	0.06%	TRIM -13%
69	RAYMOND JAMES FINANCIAL INC	38,33838,338	$924,329	0.05%	TRIM -6%
70	Cerity Partners LLC	38,18938,189	$920,737	0.05%	ADD 43%
71	CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM	36,36736,367	$876,808	0.05%	—
72	DEUTSCHE BANK AG\	36,07036,070	$869,648	0.05%	TRIM -74%
73	FOX RUN MANAGEMENT, L.L.C.	35,88735,887	$854,828	0.05%	ADD 75%
74	Creative Planning	34,07234,072	$821,477	0.05%	ADD 76%
75	ZACKS INVESTMENT MANAGEMENT	33,66333,663	$811,606	0.05%	ADD 4%
76	VANGUARD ASSET MANAGEMENT, Ltd	33,55533,555	$809,011	0.04%	NEW
77	EntryPoint Capital, LLC	33,46333,463	$806,793	0.04%	ADD 304%
78	Legal & General Group Plc	32,38532,385	$780,803	0.04%	TRIM -18%
79	TEACHER RETIREMENT SYSTEM OF TEXAS	29,52329,523	$711,799	0.04%	ADD 37%
80	AMERIPRISE FINANCIAL INC	29,25329,253	$705,289	0.04%	ADD 6%
81	SUSQUEHANNA INTERNATIONAL GROUP, LLP	27,91027,910	$672,910	0.04%	TRIM -63%
82	Universal- Beteiligungs- und Servicegesellschaft mbH	27,66327,663	$666,955	0.04%	ADD 88%
83	Atom Investors LP	25,63425,634	$618,036	0.03%	NEW
84	Fonville Wealth Management LLC	24,53324,533	$591,491	0.03%	—
85	WEALTH ENHANCEMENT ADVISORY SERVICES, LLC	22,63822,638	$553,946	0.03%	ADD 19%
86	Quantinno Capital Management LP	22,44122,441	$541,053	0.03%	NEW
87	MetLife Investment Management, LLC	22,32722,327	$538,304	0.03%	—
88	MERCER GLOBAL ADVISORS INC /ADV	20,35320,353	$490,711	0.03%	ADD 34%
89	Schonfeld Strategic Advisors LLC	19,74019,740	$475,930	0.03%	ADD 157%
90	WELLS FARGO & COMPANY/MN	19,28419,284	$464,930	0.03%	TRIM -61%
91	VOYA INVESTMENT MANAGEMENT LLC	18,96018,960	$457,126	0.03%	—
92	Zurcher Kantonalbank (Zurich Cantonalbank)	18,82718,827	$453,919	0.03%	ADD 78%
93	PRUDENTIAL FINANCIAL INC	18,01418,014	$434,318	0.02%	TRIM -93%
94	MANUFACTURERS LIFE INSURANCE COMPANY, THE	17,64217,642	$425,349	0.02%	TRIM -3%
95	Vanguard Global Advisers, LLC	16,38316,383	$394,994	0.02%	NEW
96	MARSHALL WACE, LLP	15,79215,792	$380,746	0.02%	TRIM -98%
97	SG Americas Securities, LLC	15,49415,494	$374,000	0.02%	TRIM -79%
98	SUSQUEHANNA FUNDAMENTAL INVESTMENTS, LLC	15,39715,397	$371,222	0.02%	TRIM -86%
99	JANUS HENDERSON GROUP PLC	15,16515,165	$365,477	0.02%	TRIM -66%
100	DRW Securities, LLC	15,11615,116	$364,446	0.02%	ADD 145%

† a filer this large appearing as NEW usually reflects a renamed or succeeded filing entity (the old name stops filing the same quarter) — not a brand-new position.

Rows rank by the value each filer reported — 13F filers price their own positions, so a holder with fewer shares can rank just above one with more.

Holder trends· top 15 holders · shares held over 6 quarters

green added · red trimmed

Holder

2024-Q4

2025-Q1

2025-Q2

2025-Q3

2025-Q4

2026-Q1

Latest value

VANGUARD PORTFOLIO MANAGEMENT LLC

—

4.2M

$100M

RPD Fund Management LLC

—

11K

—

3.9M

$94M

Fivespan Partners, LP

—

1.4M

3.5M

$85M

BlackRock, Inc.

3.3M

3.2M

3.4M

3.3M

3.2M

$78M

Lead Edge Capital Management, LLC

1.5M

1.6M

1.8M

1.6M

1.9M

$46M

VANGUARD CAPITAL MANAGEMENT LLC

—

1.7M

$40M

MORGAN STANLEY

691K

455K

637K

1.2M

1.4M

1.5M

$37M

ARROWSTREET CAPITAL, LIMITED PARTNERSHIP

149K

222K

691K

1.3M

1.5M

$37M

FIRST TRUST ADVISORS LP

1.3M

1.2M

1.1M

1.2M

1.4M

$33M

STATE STREET CORP

1.1M

1.0M

981K

1.0M

$25M

GEODE CAPITAL MANAGEMENT, LLC

972K

1.0M

986K

985K

$24M

AQR CAPITAL MANAGEMENT LLC

43K

200K

247K

622K

672K

958K

$23M

BAILLIE GIFFORD & CO

1.1M

1.0M

994K

941K

906K

$22M

Mudita Advisors LLP

—

839K

$20M

JACOBS LEVY EQUITY MANAGEMENT, INC

—

249K

512K

685K

621K

603K

$15M

Shares reported each quarter by the largest current holders. Green = added vs the prior filing, red = trimmed, grey = roughly held; a dash is a quarter with no 13F on file (a gap is skipped, not read as an exit). 13F is filed up to 45 days after quarter-end. Up to 40 quarters (10 years) available — use the selector above.

Institutional footprint by quarter

Line breaks are honest gaps — quarters with no 13F record, or a suppressed % — never interpolated. % of shares outstanding uses today's share count for every quarter (the only count stored).

Quarter	Holders	Shares held	% shares out	Value held
2026-Q1	206	38,913,967	52.1%	$939M
2025-Q4	246	35,834,874	48.0%	$1.3B
2025-Q3	232	36,534,417	48.9%	$1.1B
2025-Q2	221	36,160,969	48.4%	$1.1B
2025-Q1	221	35,307,504	47.3%	$1.0B
2024-Q4	204	34,276,926

Short interest has its own tab now — open SI →

Source: SEC Form 13F — every institution manager (≥ $100M in 13F securities) reporting a long US-listed position, across ~10,250 filers over 10 years of history. 13F is long-only, US-listed, filed up to 45 days after quarter-end.

Activists & large holders· Schedule 13D/G beneficial owners · APPN

Activists — 13D, active intent

Fivespan Partners, LP7.9%▲ +1.7ppSCHEDULE 13D/A · Amd 1 · event 2026-02-03SEC →

3,380,907 shares · 2 reporting persons

Abdiel Qualified Master Fund LP4.9%▼ -1.7ppSCHEDULE 13D/A · Amd 54 · event 2025-08-27SEC →

2,111,093 shares · 6 reporting persons

“Not Applicable”

Large holders — 13G, passive (top 5 of 6 by % of class)

Holder

% of class

Type

As of

Calkins Matthew W

39.2%

Exempt

2024-12-31

SEC →

VANGUARD PORTFOLIO MANAGEMENT LLC

10.06%

QII

Institutional ownership· 13F filers holding APPN · 2026-Q1 methodology → sources →

206 holders

% of shares outstanding held

52.1%

Shares held (13F)

38,913,967

Reporting holders

206

Top holders

largest 100 of 206 · Δ vs 2025-Q4

#	Holder	Shares	Value	% of shares out	Δ QoQ
1	VANGUARD PORTFOLIO MANAGEMENT LLC	4,161,5594.2M	$100M	5.57%	NEW†
2	RPD Fund Management LLC	3,919,3563.9M	$94M	5.25%	NEW†
3	Fivespan Partners, LP	3,510,9073.5M	$85M	4.70%	ADD 155%
4	BlackRock, Inc.	3,226,0873.2M	$78M	4.32%	—
5	Lead Edge Capital Management, LLC	1,905,6341.9M	$46M	2.55%	ADD 18%
6	VANGUARD CAPITAL MANAGEMENT LLC	1,678,2801.7M	$40M	2.25%	NEW†
7	MORGAN STANLEY	1,545,8631.5M	$37M	2.07%	ADD 9%
8	ARROWSTREET CAPITAL, LIMITED PARTNERSHIP	1,519,6531.5M	$37M	2.04%	ADD 16%
9	FIRST TRUST ADVISORS LP	1,355,9611.4M	$33M	1.82%	ADD 17%
10	STATE STREET CORP	1,023,4421.0M	$25M	1.37%	ADD 4%
11	GEODE CAPITAL MANAGEMENT, LLC	985,499985,499	$24M	1.32%	—
12	AQR CAPITAL MANAGEMENT LLC	957,620957,620	$23M	1.28%	ADD 43%
13	BAILLIE GIFFORD & CO	905,908905,908	$22M	1.21%	TRIM -4%
14	Mudita Advisors LLP	839,400839,400	$20M	1.12%	NEW
15	JACOBS LEVY EQUITY MANAGEMENT, INC	603,430603,430	$15M	0.81%	TRIM -3%
16	ACADIAN ASSET MANAGEMENT LLC	520,419520,419	$13M	0.70%	TRIM -15%
17	CITADEL ADVISORS LLC	518,009518,009	$12M	0.69%	TRIM -6%
18	D. E. Shaw & Co., Inc.	447,788447,788	$11M	0.60%	TRIM -60%
19	Nuveen, LLC	414,956414,956	$10M	0.56%	TRIM -40%
20	Squarepoint Ops LLC	390,929390,929	$9M	0.52%	ADD 32%
21	PRELUDE CAPITAL MANAGEMENT, LLC	390,321390,321	$9M	0.52%	NEW
22	FMR LLC	380,697380,697	$9M	0.51%	ADD 18%
23	CHARLES SCHWAB INVESTMENT MANAGEMENT INC	370,662370,662	$9M	0.50%	ADD 6%
24	MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.	353,022353,022	$9M	0.47%	TRIM -33%
25	NORTHERN TRUST CORP	344,336344,336	$8M	0.46%	ADD 2%
26	Trexquant Investment LP	305,943305,943	$7M	0.41%	ADD 97%
27	National Philanthropic Trust	299,532299,532	$7M	0.40%	—
28	Qube Research & Technologies Ltd	292,737292,737	$7M	0.39%	TRIM -39%
29	GOLDMAN SACHS GROUP INC	285,265285,265	$7M	0.38%	TRIM -30%
30	UBS Group AG	284,284284,284	$7M	0.38%	ADD 205%
31	VANGUARD FIDUCIARY TRUST CO	277,406277,406	$7M	0.37%	NEW
32	Connor, Clark & Lunn Investment Management Ltd.	208,561208,561	$5M	0.28%	TRIM -43%
33	JANE STREET GROUP, LLC	194,452194,452	$5M	0.26%	TRIM -23%
34	HRT FINANCIAL LP	189,984189,984	$5M	0.25%	ADD 79%
35	ALGERT GLOBAL LLC	179,554179,554	$4M	0.24%	ADD 23%
36	CANADA PENSION PLAN INVESTMENT BOARD	179,500179,500	$4M	0.24%	TRIM -9%
37	BANK OF AMERICA CORP /DE/	176,566176,566	$4M	0.24%	ADD 8%
38	Tejara Capital Ltd	173,300173,300	$4M	0.23%	NEW
39	Harber Asset Management LLC	147,426147,426	$4M	0.20%	TRIM -22%
40	Point72 Asset Management, L.P.	147,228147,228	$4M	0.20%	TRIM -36%
41	ExodusPoint Capital Management, LP	146,023146,023	$4M	0.20%	NEW
42	JPMORGAN CHASE & CO	145,224145,224	$3M	0.19%	ADD 30%
43	Engineers Gate Manager LP	130,979130,979	$3M	0.18%	TRIM -15%
44	BNP PARIBAS FINANCIAL MARKETS	127,268127,268	$3M	0.17%	TRIM -48%
45	Russell Investments Group, Ltd.	123,051123,051	$3M	0.16%	TRIM -49%
46	Bank of New York Mellon Corp	120,429120,429	$3M	0.16%	TRIM -2%
47	TWO SIGMA INVESTMENTS, LP	119,066119,066	$3M	0.16%	TRIM -63%
48	Gotham Asset Management, LLC	95,64895,648	$2M	0.13%	ADD 99%
49	UBS AM, a distinct business unit of UBS ASSET MANAGEMENT AMERICAS LLC	86,00086,000	$2M	0.12%	ADD 2%
50	SEI INVESTMENTS CO	85,09985,099	$2M	0.11%	ADD 8%
51	LOS ANGELES CAPITAL MANAGEMENT LLC	80,62480,624	$2M	0.11%	ADD 107%
52	Swiss National Bank	76,30076,300	$2M	0.10%	—
53	BOOTHBAY FUND MANAGEMENT, LLC	75,16075,160	$2M	0.10%	NEW
54	BARCLAYS PLC	74,74874,748	$2M	0.10%	TRIM -44%
55	ALLIANCEBERNSTEIN L.P.	44,56144,561	$2M	0.06%	ADD 5%
56	IEQ CAPITAL, LLC	63,24863,248	$2M	0.08%	TRIM -24%
57	Brevan Howard Capital Management LP	58,89858,898	$1M	0.08%	ADD 495%
58	CWM, LLC	53,82053,820	$1M	0.07%	ADD 65%
59	RHUMBLINE ADVISERS	51,62751,627	$1M	0.07%	TRIM -25%
60	PRICE T ROWE ASSOCIATES INC /MD/	49,22649,226	$1M	0.07%	ADD 18%
61	Magnetar Financial LLC	48,07148,071	$1M	0.06%	ADD 81%
62	FRANKLIN RESOURCES INC	47,23847,238	$1M	0.06%	ADD 186%
63	PDT Partners, LLC	47,08947,089	$1M	0.06%	TRIM -58%
64	Invesco Ltd.	46,98446,984	$1M	0.06%	—
65	CITIGROUP INC	45,91945,919	$1M	0.06%	TRIM -79%
66	Mariner, LLC	42,87442,874	$1M	0.06%	ADD 78%
67	Amundi	42,80442,804	$1M	0.06%	TRIM -67%
68	Man Group plc	42,44042,440	$1M	0.06%	TRIM -13%
69	RAYMOND JAMES FINANCIAL INC	38,33838,338	$924,329	0.05%	TRIM -6%
70	Cerity Partners LLC	38,18938,189	$920,737	0.05%	ADD 43%
71	CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM	36,36736,367	$876,808	0.05%	—
72	DEUTSCHE BANK AG\	36,07036,070	$869,648	0.05%	TRIM -74%
73	FOX RUN MANAGEMENT, L.L.C.	35,88735,887	$854,828	0.05%	ADD 75%
74	Creative Planning	34,07234,072	$821,477	0.05%	ADD 76%
75	ZACKS INVESTMENT MANAGEMENT	33,66333,663	$811,606	0.05%	ADD 4%
76	VANGUARD ASSET MANAGEMENT, Ltd	33,55533,555	$809,011	0.04%	NEW
77	EntryPoint Capital, LLC	33,46333,463	$806,793	0.04%	ADD 304%
78	Legal & General Group Plc	32,38532,385	$780,803	0.04%	TRIM -18%
79	TEACHER RETIREMENT SYSTEM OF TEXAS	29,52329,523	$711,799	0.04%	ADD 37%
80	AMERIPRISE FINANCIAL INC	29,25329,253	$705,289	0.04%	ADD 6%
81	SUSQUEHANNA INTERNATIONAL GROUP, LLP	27,91027,910	$672,910	0.04%	TRIM -63%
82	Universal- Beteiligungs- und Servicegesellschaft mbH	27,66327,663	$666,955	0.04%	ADD 88%
83	Atom Investors LP	25,63425,634	$618,036	0.03%	NEW
84	Fonville Wealth Management LLC	24,53324,533	$591,491	0.03%	—
85	WEALTH ENHANCEMENT ADVISORY SERVICES, LLC	22,63822,638	$553,946	0.03%	ADD 19%
86	Quantinno Capital Management LP	22,44122,441	$541,053	0.03%	NEW
87	MetLife Investment Management, LLC	22,32722,327	$538,304	0.03%	—
88	MERCER GLOBAL ADVISORS INC /ADV	20,35320,353	$490,711	0.03%	ADD 34%
89	Schonfeld Strategic Advisors LLC	19,74019,740	$475,930	0.03%	ADD 157%
90	WELLS FARGO & COMPANY/MN	19,28419,284	$464,930	0.03%	TRIM -61%
91	VOYA INVESTMENT MANAGEMENT LLC	18,96018,960	$457,126	0.03%	—
92	Zurcher Kantonalbank (Zurich Cantonalbank)	18,82718,827	$453,919	0.03%	ADD 78%
93	PRUDENTIAL FINANCIAL INC	18,01418,014	$434,318	0.02%	TRIM -93%
94	MANUFACTURERS LIFE INSURANCE COMPANY, THE	17,64217,642	$425,349	0.02%	TRIM -3%
95	Vanguard Global Advisers, LLC	16,38316,383	$394,994	0.02%	NEW
96	MARSHALL WACE, LLP	15,79215,792	$380,746	0.02%	TRIM -98%
97	SG Americas Securities, LLC	15,49415,494	$374,000	0.02%	TRIM -79%
98	SUSQUEHANNA FUNDAMENTAL INVESTMENTS, LLC	15,39715,397	$371,222	0.02%	TRIM -86%
99	JANUS HENDERSON GROUP PLC	15,16515,165	$365,477	0.02%	TRIM -66%
100	DRW Securities, LLC	15,11615,116	$364,446	0.02%	ADD 145%

† a filer this large appearing as NEW usually reflects a renamed or succeeded filing entity (the old name stops filing the same quarter) — not a brand-new position.

Rows rank by the value each filer reported — 13F filers price their own positions, so a holder with fewer shares can rank just above one with more.

Holder trends· top 15 holders · shares held over 6 quarters

green added · red trimmed

Holder

2024-Q4

2025-Q1

2025-Q2

2025-Q3

2025-Q4

2026-Q1

Latest value

VANGUARD PORTFOLIO MANAGEMENT LLC

—

4.2M

$100M

RPD Fund Management LLC

—

11K

—

3.9M

$94M

Fivespan Partners, LP

—

1.4M

3.5M

$85M

BlackRock, Inc.

3.3M

3.2M

3.4M

3.3M

3.2M

$78M

Lead Edge Capital Management, LLC

1.5M

1.6M

1.8M

1.6M

1.9M

$46M

VANGUARD CAPITAL MANAGEMENT LLC

—

1.7M

$40M

MORGAN STANLEY

691K

455K

637K

1.2M

1.4M

1.5M

$37M

ARROWSTREET CAPITAL, LIMITED PARTNERSHIP

149K

222K

691K

1.3M

1.5M

$37M

FIRST TRUST ADVISORS LP

1.3M

1.2M

1.1M

1.2M

1.4M

$33M

STATE STREET CORP

1.1M

1.0M

981K

1.0M

$25M

GEODE CAPITAL MANAGEMENT, LLC

972K

1.0M

986K

985K

$24M

AQR CAPITAL MANAGEMENT LLC

43K

200K

247K

622K

672K

958K

$23M

BAILLIE GIFFORD & CO

1.1M

1.0M

994K

941K

906K

$22M

Mudita Advisors LLP

—

839K

$20M

JACOBS LEVY EQUITY MANAGEMENT, INC

—

249K

512K

685K

621K

603K

$15M

Institutional footprint by quarter

Line breaks are honest gaps — quarters with no 13F record, or a suppressed % — never interpolated. % of shares outstanding uses today's share count for every quarter (the only count stored).

Quarter	Holders	Shares held	% shares out	Value held
2026-Q1	206	38,913,967	52.1%	$939M
2025-Q4	246	35,834,874	48.0%	$1.3B
2025-Q3	232	36,534,417	48.9%	$1.1B
2025-Q2	221	36,160,969	48.4%	$1.1B
2025-Q1	221	35,307,504	47.3%	$1.0B
2024-Q4	204	34,276,926

Short interest has its own tab now — open SI →

Activists & large holders· Schedule 13D/G beneficial owners · APPN

Activists — 13D, active intent

Fivespan Partners, LP7.9%▲ +1.7ppSCHEDULE 13D/A · Amd 1 · event 2026-02-03SEC →

3,380,907 shares · 2 reporting persons

Abdiel Qualified Master Fund LP4.9%▼ -1.7ppSCHEDULE 13D/A · Amd 54 · event 2025-08-27SEC →

2,111,093 shares · 6 reporting persons

“Not Applicable”

Large holders — 13G, passive (top 5 of 6 by % of class)

Holder

% of class

Type

As of

Calkins Matthew W

39.2%

Exempt

2024-12-31

SEC →

VANGUARD PORTFOLIO MANAGEMENT LLC

10.06%

QII

APPN US Equity☆

Institutional ownership· 13F filers holding APPN · 2026-Q1 methodology → sources →

Top holders

Holder trends· top 15 holders · shares held over 6 quarters

Institutional footprint by quarter

Activists & large holders· Schedule 13D/G beneficial owners · APPN

Institutional ownership· 13F filers holding APPN · 2026-Q1 methodology → sources →

Top holders

Holder trends· top 15 holders · shares held over 6 quarters

Institutional footprint by quarter

Activists & large holders· Schedule 13D/G beneficial owners · APPN

APPN US Equity